Colymboides

Colymboides; Temporal range:
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Class:	Aves
Order:	Gaviiformes
Family:	†Colymboididae; Brodkorb, 1963
Subfamily:	†Colymboidinae; Brodkorb, 1963
Genus:	†Colymboides; Milne-Edwards, 1867
Species
	†C. minutus Milne-Edwards, 1867; †C. anglicus Lydekker, 1891; †C. metzleri Mayr, 2004[1]
Synonyms
	Gavioides

The genus Colymboides contains three species of early loon dating from the late Oligocene or early Miocene. They are considered to be the earliest known unambiguous gaviiform fossils. The genus is widely known from early Priabonian – about 37 million years ago in the Late Eocene – to Early Miocene (late Burdigalian, less than 20 million years ago) limnic and marine rocks of western Eurasia north of the Alpide belt, between the Atlantic and the former Turgai Sea. It is usually placed in the Gaviidae already, but usually[Note 1] in a subfamily Colymboidinae, with the modern-type loons making up the Gaviinae. But the Colymboides material is generally quite distinct from modern loons, and may actually belong in a now-extinct family of primitive gaviiforms. The best studied species, Colymboides minutus, was described by Robert Storer[2] as being much smaller than modern loons and not as well adapted to diving.

Notes

Some (notably Robert W. Storer) have disagreed, usually because they separated Gaviella in the basalmost subfamily of the Gaviidae and considered Colymboides the ancestor of Gavia. More recent authors generally disagree at least regarding the latter: Storer (1956),[2] Olson (1985),[3] Mayr (2009)[4]

References

Mayr G (2004). "A partial skeleton of a new fossil loon (Aves, Gaviiformes) from the early Oligocene of Germany with preserved stomach content". Journal of Ornithology. 145 (4): 281–286. doi:10.1007/s10336-004-0050-9. S2CID 1070943.
Robert W. Storer (1956). "The fossil loon, Colymboides minutus". The Condor. 58 (6): 413–426. doi:10.2307/1365096. JSTOR 1365096.
Storrs Olson (1985). "Section X.I. Gaviiformes" (PDF). In D. S. Farner, J. R. King & Kenneth C. Parkes (ed.). Avian Biology. Vol. 8. pp. 212–214.
Mayr, G. (2009). "Gaviiformes". In Gerald Mayr (ed.). Paleogene Fossil Birds. Heidelberg & New York: Springer Verlag. pp. 75–76. doi:10.1007/978-3-540-89628-9_7. ISBN 978-3-540-89627-2.

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[5] Some (notably Robert W. Storer) have disagreed, usually because they separated Gaviella in the basalmost subfamily of the Gaviidae and considered Colymboides the ancestor of Gavia. More recent authors generally disagree at least regarding the latter: Storer (1956),[2] Olson (1985),[3] Mayr (2009)[4]

[1] Mayr G (2004). "A partial skeleton of a new fossil loon (Aves, Gaviiformes) from the early Oligocene of Germany with preserved stomach content". Journal of Ornithology. 145 (4): 281–286. doi:10.1007/s10336-004-0050-9. S2CID 1070943.

[Storer-2] Robert W. Storer (1956). "The fossil loon, Colymboides minutus". The Condor. 58 (6): 413–426. doi:10.2307/1365096. JSTOR 1365096.

[3] Storrs Olson (1985). "Section X.I. Gaviiformes" (PDF). In D. S. Farner, J. R. King & Kenneth C. Parkes (ed.). Avian Biology. Vol. 8. pp. 212–214.

[4] Mayr, G. (2009). "Gaviiformes". In Gerald Mayr (ed.). Paleogene Fossil Birds. Heidelberg & New York: Springer Verlag. pp. 75–76. doi:10.1007/978-3-540-89628-9_7. ISBN 978-3-540-89627-2.