Alcobaça Formation
The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit,[1] is a geological formation in Portugal.[2] It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of information on the diversity of Late Jurassic mammals.[3][4] Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.
| Alcobaça Formation | |
|---|---|
| Stratigraphic range: Kimmeridgian ~ | |
| Type | Geological formation |
| Sub-units | Camadas de Guimarota |
| Underlies | Lourinhã Formation |
| Overlies | Montejunto Formation |
| Lithology | |
| Primary | Marl |
| Other | Mudstone, coal, siltstone, limestone, sandstone |
| Location | |
| Coordinates | 39.6°N 9.0°W |
| Approximate paleocoordinates | 33.7°N 1.6°E |
| Region | Estremadura, Leiria, Lisboa |
| Country |  Portugal |
| Extent | Lusitanian Basin |
| Type section | |
| Named for | Alcobaça, Portugal |
| Named by | Choffat |
| Year defined | 1885 |
  Alcobaça Formation (Portugal) | |
Vertebrate paleofauna
Dinosaur eggs are geographically located in Lisbon District, Portugal.[5] Dinosaur tracks are geographically located in Leiria District, Portugal.[6]
Amphibians
| Amphibians of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
| Celtedens | Indeterminate | Porto Pinheiro | Albanerpetontid | |||
| Choristoderes of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
| Cteniogenys | Indeterminate | Guimarota | ||||
Ornithischians
Indeterminate euornithopod remains located in Lisbon District.[5] Indeterminate stegosaurid remains present in Lisbon District.[5]
| Ornithischians of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | District | Stratigraphic position | Abundance | Notes | Images |
| Dacentrurus[5] | D. armatus[5] | Lisboa[5] |  | |||
| Ornithopoda[5] | Indeterminate[5] | Lisboa[5] | ||||
| Stegosaurus[7] | S. cf. ungulatus | "...one tooth, five cervical vertebrae (including the axis), five dorsal vertebrae, cervical and dorsal ribs, three caudal vertebrae, chevrons, preacetabular process of the left ilium, right tibia, fibula, astragalus and [calcaneus], a cervical plate, and several fragments of plates." |  | |||
| Trimucrodon[5] | T. cuneatus[5] | Lisboa[5] | Later determined to be indeterminate ornithischian remains.[5] | |||
Saurischians
Indeterminate sauropod remains located in Leiria and Lisboa.[8]
| Saurischians of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | District | Stratigraphic position | Abundance | Notes | Images |
| Allosaurus[6] | Indeterminate[6] | Leiria[6] |  | |||
| Aviatyrannis | A. jurassica | Guimarota | ||||
| Lusotitan[5] | L. atalaiensis[5] | Lisboa[5] |  | |||
| Ceratosaurus[5] | Indeterminate[5] | Lisboa[5] |  | |||
| Dinheirosaurus[5] | D. lourinhanensis[5] | Lisboa[5] | "[Two] cervicals, [nine] dorsals, ribs, limb elements."[9] |  | ||
| Lourinhasaurus[5] | L. alenquerensis[5] | Lisboa[5] | ||||
| Megalosaurus[5] | M. insignis[5] | Lisboa[5] | Later determined to be indeterminate theropod remains.[5] | |||
Turtles
| Turtles of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
| Peltochelys | P. duchastelli | Guimarota | ||||
| Platychelys | Indeterminate | Guimarota | ||||
| Pleurosternidae | Indeterminate | Guimarota | ||||
Lepidosauromorphs
| Lepidosauromorphs of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
| Becklesius | B. hoffstetteri | Guimarota | ||||
| Dorsetisaurus | D. purbeckensis | Guimarota | ||||
| Marmoretta | Indeterminate | Guimarota | Basal Lepidosauromorph | |||
| Portugalophis | P. lignites | Guimarota | Stem-snake | |||
| Saurillodon | S. proraformis | Guimarota | ||||
| Saurillus | S. henkeli, cf. obtusus | Guimarota | ||||
Crocodyliformes
| Crocodyliformes of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
| Goniopholis | G. baryglyphaeus | Guimarota |  | |||
| Knoetschkesuchus | K. guimarotae | Guimarota |  | |||
| Lisboasaurus | L. mitracostatus | Guimarota | ||||
| Machimosaurus | M. hugii | Guimarota |  | |||
Mammals
| Mammals of the Alcobaça Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
| Bathmochoffatia | B. hapax | Guimarota | A paulchoffatiid multituberculate | |||
| Bernardodon | B. sp, B. atlanticus | Porto Pinheiro | A pinheirodontid multituberculate | |||
| Drescheratherium | D. acutum | Guimarota | A paurodontid dryolestidan | |||
| Dryolestes | D. leirensis | Guimarota | A dryolestid dryolestidan | |||
| Ecprepaulax | E. anomala | Porto Pinheiro | A pinheirodontid multituberculate | |||
| Guimarotodon | G. leiriensis | Guimarota | A paulchoffatiid multituberculate | |||
| Guimarotodus | G. inflatus | Guimarota | A dryolestid dryolestidan | |||
| Haldanodon | H. exspectatus | Guimarota | A docodont | |||
| Henkelodon | H. naias, H. sp | Guimarota | A paulchoffatiid multituberculate | |||
| Henkelotherium | H. guimarotae | Guimarota | A dryolestidan | |||
| Iberodon | I. quadrituberculatus | Porto Pinheiro | A pinheirodontid multituberculate | |||
| Kielanodon | K. hopsoni | Guimarota | A paulchoffatiid multituberculate | |||
| Krebsotherium | K. lusitanicum | Guimarota | A dryolestid dryolestidan | |||
| Kuehneodon | K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp | Guimarota | A paulchoffatiid multituberculate | |||
| Laolestes | L. andresi | Porto Pinheiro | A dryolestid dryolestidan | |||
| Meketibolodon | M. robustus | Guimarota | A paulchoffatiid multituberculate | |||
| Meketichoffatia | M. krausei, M. sp | Guimarota | A paulchoffatiid multituberculate | |||
| Nanolestes | N. krusati | Porto Pinheiro | A cladotherian | |||
| N. drescherae | Guimarota | |||||
| Paulchoffatia | P. delgadoi | Guimarota | A paulchoffatiid multituberculate | |||
| Pinheirodon | P. sp, P. vastus, P. pygmaeus | Porto Pinheiro | A pinheirodontid multituberculate | |||
| Plesiochoffatia | P. thoas, P. peperethos | Guimarota | A paulchoffatiid multituberculate | |||
| Portopinheirodon | P. asymmetricus | Porto Pinheiro | A dryolestid dryolestidan | |||
| Priacodon | Indeterminate | Porto Pinheiro | A triconodontid | |||
| Proalbionbaatar | P. plagiocyrtus | Guimarota | A multituberculate of uncertain placement | |||
| Pseudobolodon | P. dryas, P krebsi, P. oreas | Guimarota | A paulchoffatiid multituberculate | |||
| Renatodon | R. amalthea | Guimarota | A paulchoffatiid multituberculate | |||
| ? Tinodon | Indeterminate | Porto Pinheiro | A tinodontid symmetrodont | |||
| Xenachoffatia | X. oinopion | Guimarota | A paulchoffatiid multituberculate | |||
| Paulchoffatiidae | Indeterminate | Guimarota | 4 distinct genera with 7 species | |||
| Pinheirodontidae | Indeterminate | Porto Pinheiro | ||||
Color key
|
Notes Uncertain or tentative taxa are in small text; |
See also
References
- Mateus, O.; Dinis, J.; Cunha, P. P. (2017-09-28). "The Lourinhã Formation: the Upper Jurassic to lower most Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked". Ciências da Terra / Earth Sciences Journal. 19 (1): 75–97. doi:10.21695/cterra/esj.v19i1.355. ISSN 2183-4431.
- Fürsich, Franz T.; Schneider, Simon; Werner, Winfried; Lopez-Mir, Berta; Pierce, Colm S. (2021-09-13). "Life at the continental–marine interface: palaeoenvironments and biota of the Alcobaça Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended". Palaeobiodiversity and Palaeoenvironments. doi:10.1007/s12549-021-00496-x. ISSN 1867-1608.
- "Guimarota". Paleobiology Database. Retrieved 31 August 2018.
- "Porto Pinheiro, Lourinha". Paleobiology Database. Retrieved 31 August 2018.
- "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
- "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
- Escaso, Fernando; Ortega, Francisco; Dantas, Pedro; Malafaia, Elisabete; Pimentel, Nuno L.; Pereda-Suberbiola, Xabier; Sanz, José Luis; Kullberg, José Carlos; Kullberg, María Carla (2006-12-23). "New evidence of shared dinosaur across Upper Jurassic Proto-North Atlantic: Stegosaurus from Portugal". Naturwissenschaften. 94 (5): 367–374. Bibcode:2007NW.....94..367E. doi:10.1007/s00114-006-0209-8. PMID 17187254. S2CID 10930309.
- "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
- "Table 13.1," in Weishampel, et al. (2004). Page 265.
Bibliography
- Weishampel, David B.; Peter Dodson, and Halszka (eds.) Osmólska. 2004. The Dinosauria, 2nd edition, 1–880. Berkeley: University of California Press. Accessed 2019-02-21. ISBN 0-520-24209-2
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