Humulus japonicus

Humulus japonicus (syn. H. scandens, Antidesma scandens) is known by the common name of "Japanese hop",[4][5] or "wild hop".[6]

The plant is called lü cao (葎草) in Mandarin Chinese, and kanamugura (カナムグラ) in Japanese.[7]

Leaves

Flower

Well-developed leaves with 5-9 lobes; abaxial surfaces of leaves pubescent, but not densely so; infructescence rarely longer than 2 cm, with bracts and bracteoles less than 1 cm long, and notable spinulose-ciliate[8] Inflorescences: staminate inflorescences erect, 15–25 cm, flower anthers without glands such as; pistillate inflorescences spikes, conelike, ovoid; bracteole ovate-orbiculate, 7–10 mm, pilose, margins densely ciliate-hairy. Infructescences pendulous, green, conelike, ovoid to oblong, (1–)1.5–3(–4) cm; bracteoles without yellow glands.[9]

Seeds germinate in early spring, but plants may continue to emerge if conditions are favorable. Seeds can be dispersed by number of agents including humans, animals, machines, and flood waters.[10]

Humulus japonicus is native temperate parts of Asia (China, Japan, Korea, Taiwan and the Russian Far East) and the tropical environment of Vietnam and Laos,[10][4] but has become an invasive species in North America since it was imported in the late 19th century.[11]

The plant has been immensely successful in North America, and is found throughout eastern Canada and the eastern half of the United States, and considered a invasive species.[12] In the United States Humulus japonicus has a lateral range from Nebraska/North Dakota to Maine and a vertical range from Minnesota to Georgia.[10][13]

Japanese hops grow well given plentiful sunlight, moisture, and nutrient rich soil, and are most commonly found along stream banks and floodplains.[10] Japanese hops do not grow as well in shaded areas and drier soils.[10] In milder climates, it can survive the winter.[10]

The plant was introduced into America in the 19th century both as an ornamental and raw ingredient for the Asian tonic.[10] In traditional Chinese medicine this herb known as lü cao is used rather uncommonly, with supposed diuretic and heat-cooling effects.[14][15]

Although related to the common hop Humulus lupulus, whose cultivars are the brewer's hop used in beer production, the Japanese hop is found to be deficient in the bitter acids and essential oils used for flavoring beer, being devoid of lupulin glands (or producing very few resin glands[16]); hence the Japanese hop is not considered viable for use in brewing.[10][17][11]

This species is a host plant for the butterfly Polygonia c-aureum, the Asian comma, hence in China the butterfly is considered a major pest affecting the cash crop used in medicine.[18]

The Japanese hop can adapt well to new environments. Its range and distribution in North America prove that it can thrive in a wide range of environments, from the southern regions of the U.S. to the far reaches of Northern Canada. It can grow and cover any area quickly. Thousands of hop plants can grow in one acre, and they can blanket a majority of the local vegetation.[10] The vines grow rapidly during the summer, climbing up and over everything in their path, and can form dense mats several feet deep, blocking light to whatever plants are underneath.[10]

Citations

Wikimedia Commons has media related to Humulus japonicus.

• invasive.org – Japanese hop
• botanic.jp – some photos