Ichneutica cuneata

Ichneutica cuneata is a moth in the family Noctuidae.[1] It is endemic to New Zealand. I. cuneata is found in and around the Tongariro National Park in the North Island and throughout the South Island. Specimens found in Tongariro National Park tend to be darker in appearance than South Island specimens but as their colour is variable, and antennae and genitalia of both species are consistent, they are not now regarded as a separate species. I. cuneata inhabits tussock grasslands and shrublands in alpine and subalpine zones. The adults are on the wing from December to April and can be found flying during the day. The life history of I. cuneata is unknown as are the specific host species of its larvae. However larvae of I. cuneata are said to feed on herbaceous plants. The adults of this species also pollinate native species such as Myosotis macrantha.

Ichneutica cuneata
Female
Male
Scientific classification
Kingdom:
Phylum:
Class:
Order:
Family:
Genus:
Species:
I. cuneata
Binomial name
Ichneutica cuneata
(Philpott, 1916)[1]
Synonyms
  • Aletia cuneata Philpott,1916
  • Aletia s.l. cuneata (Philpott, 1916)
  • Aletia dentata Philpott, 1923
  • Mythimna cuneata (Philpott, 1916)
  • Mythimna dentata (Philpott, 1923)
  • Aletia s.l. dentata (Philpott, 1923)

Taxonomy

The species was first described by Alfred Philpott in 1916 from specimens obtained at Ben Lomond and Macetown.[2] Philpott named the species Aletia cuneata.[2][3] The lectotype specimen, collected at Ben Lomond in January, is held in the New Zealand Arthropod Collection.[4] In 1988 J. S. Dugdale, in his catalogue on New Zealand Lepidopera, placed this species within the Aletia genus.[5] In 2019 Robert Hoare undertook a major review of New Zealand Noctuidae species.[4] During this review the genus Ichneutica was greatly expanded and the genus Aletia was subsumed into that genus as a synonym.[4] As a result of this review, this species is now known as Ichneutica cuneata.[4] Hoare, while undertaking the review, also inspected the type material of Aletia dentata.[4] Hoare placed this species within the genus Ichneutica and subsumed it as a synonym of Ichneutica cuneata.[4] Hoare justified this action by explaining that although the moths formerly regarded as a separate species tend to be darker, this colouration is variable and the lighter coloured specimens are identical to I. cuneata.[4] Hoare also points out that any differences in genitalia or antennae are not consistent enough to support a separate species.[4]

Description

Ichneutica cuneata observed in Cromwell
Female darker form collected at Tongariro National Park

Philpott reported that the species had a 38 to 39 mm wingspan.[2] George Hudson described the species as follows:

The forewings are greyish-ochreous, strongly clouded with brownish-ochreous in the disc; all lines are very obscure, the orbicular is almost round, outlined in black towards the termen; the claviform very small with cloudy blackish edging; the reniform dull black; there is a cloudy median shade; all the leading veins are broadly marked in blueish-gray; a blackish patch is situated near the middle of the subterminal area, two at the cornus and another on vein 1 at the base of the wing. The hind-wings are greyish-ochreous with a very broad blackish terminal band.[6]

The adult male of this species has a wingspan of between 32 and 39 mm where as the female has a wingspan of between 34 and 39 mm.[4] I. cuneata can be distinguished from similar species by the black to dark grey kidney mark on the forewings of the species as well as the colour of the hindwing which gradually changes from ocherous-grey to a darker grey along the border.[4] The underside of the hind wing also has a discal spot as well as a line. No similar species has these markings.[4]

The population of this species found at Tongariro National Park and previously separated into a distinct species tends to be darker in appearance as a result of the blackish markings on the forewings. However these markings are variable.[4]

Distribution

This species is endemic to New Zealand.[7] This species is only known from the Tongariro National Park in the North Island but is widespread in the South Island.[4] I. cuneata was discovered by J. H. Lewis at Ophir.[6] Along with Otago,[8] I. cuneata has also been found in Canterbury in the South Island,[9] as well as Mount Tongariro and Mount Ruapehu in the North Island.[6]

Habitat

This species frequents tussock grassland and shrubland in montane and subalpine areas.[4][10] They have been recorded at altitudes of between 450 and 1640 m.[11]

Behaviour

The adult moths are on the wing from December to April and can be active during the day.[4][10] They have been known to rest of rocks.[4]

Life history and host plants

Myosotis macrantha, a plant known to host and be pollinated by adult I. cuneata moths

The life history of this species is unknown as are the specific host species of its larvae.[4] However larvae of I. cuneata are said to feed on herbs.[12][13] Adults have been shown to feed from and help pollinate Myosotis macrantha at Mount Cook.[9]

References

  1. "Ichneutica cuneata (Philpott, 1916)". www.nzor.org.nz. Retrieved 2021-08-03.
  2. Philpott, Alfred (1916). "Descriptions of new species of Lepidoptera". Transactions and Proceedings of the New Zealand Institute. 48: 420–423 via Biodiversity Heritage Library.
  3. Dugdale, John S. (1988). "Lepidoptera - annotated catalogue, and keys to family-group taxa" (PDF). Fauna of New Zealand. 14: 199. ISBN 0-477-02518-8. Archived from the original (PDF) on 2019-01-27. Retrieved 2018-07-14.
  4. Hoare, Robert J. B. (9 December 2019). "Noctuinae (Insecta: Lepidoptera: Noctuidae) part 2: Nivetica, Ichneutica". Fauna of New Zealand. 80: 1–455. doi:10.7931/J2/FNZ.80.
  5. Dugdale, John Stewart (1988-09-23). "Lepidoptera - annotated catalogue, and keys to family-group taxa" (PDF). Fauna of New Zealand. 14: 1–264.
  6. Hudson, G. V. (1928). The Butterflies and Moths of New Zealand. Wellington: Ferguson & Osborn Ltd. pp. 56–57. Archived from the original on 2020-09-28. Retrieved 2017-12-18.
  7. "Ichneutica cuneata (Philpott, 1916)". nztcs.org.nz. 2021. Retrieved 2021-08-04.
  8. Patrick, Brian (1994). "Hawkdun Ecological District Invertebrate Survey" (PDF). Science & Research Series. 64. ISBN 0-478-01553-4. ISSN 0113-3713. Archived from the original (PDF) on 2017-05-07. Retrieved 2017-12-18.
  9. Newstrom, Linda; Robertson, Alastair (1 January 2005). "Progress in understanding pollination systems in New Zealand". New Zealand Journal of Botany. 43 (1): 1–59. doi:10.1080/0028825x.2005.9512943. ISSN 0028-825X. S2CID 83798157.
  10. Barratt, B. I. P.; Patrick, B. H. (1 January 1987). "Insects of snow tussock grassland on the East Otago Plateau". New Zealand Entomologist. 10 (1): 69–98. doi:10.1080/00779962.1987.9722513. ISSN 0077-9962.
  11. Patrick, Brian H.; Lyford, Brian M.; Ward, John B.; Barratt, Barbara I. P. (1 December 1992). "Lepidoptera and other insects of the Rastus Burn Basin, The Remarkables, Otago". Journal of the Royal Society of New Zealand. 22 (4): 265–278. doi:10.1080/03036758.1992.10420820. ISSN 0303-6758.
  12. White, E.G. (1991). "The changing abundance of moths in a tussock grassland, 1962-1989, and 50- to 70-year trends". New Zealand Journal of Ecology. 15 (1): 5–22. S2CID 89398506.
  13. Patrick, B.H. (1991). Insects of the Dansey ecological district (PDF). Wellington, N.Z.: Department of Conservation, New Zealand. ISBN 0-478-01285-3. Retrieved 18 December 2017.
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