Lilian Vaughan Morgan

Morgan was born in 1870 in Hallowell, Maine. She was orphaned at the age of three when her parents and younger sister died of tuberculosis. After the death of her parents, Morgan and her older sister Edith were raised by her maternal grandparents in Germantown, Pennsylvania.

Morgan enrolled as an undergraduate student at Bryn Mawr in 1887. She majored in biology and was advised by Martha Carey Thomas. After graduating with honors in 1891, she spent the summer at the Marine Biological Laboratory in Woods Hole, Massachusetts, where Edmund Beecher Wilson, one of her previous zoology professors, introduced her to her future graduate advisor and husband, Thomas Hunt Morgan.[1]

In the autumn of 1891, a Morgan earned a fellowship, which enabled her to study the musculature of chitons at the University of Zurich with Arnold Lang, a comparative anatomist and student of Ernst Haeckel.[1] She returned to Bryn Mawr in 1892, where she earned a Master of Science in biology in 1894, under the advisory of Thomas Morgan. After graduation, she published her work on the musculature of chitons, returned to Woods Hole as an independent investigator, and spent many summers investigating breeding, development and embryology in amphibians.[1]

In 1904, at the age of 34, she married Thomas Hunt Morgan and moved to New York City, where Thomas Morgan began working at Columbia University. They spent the following summer in California, where she researched and published work on planarian regeneration at the Stanford Marine Laboratory. She would not publish another paper for sixteen years.

During the years following her marriage, she supported her husband's career and raised four children: Howard Key Morgan, born 1906; Edith Sampson Morgan, born 1907; Lilian Vaughan Morgan, born 1910; and Isabella Merrick Morgan, born 1911. Shine and Wrobel (1976) note that one key to Thomas Hunt Morgan's success was that his personal affairs were entirely handled by Lilian Morgan, freeing him to focus on his research.[2] The family spent their winters in New York and returned in the summers to Woods Hole, where she maintained a summer house for her children, relatives and her husband's graduate students. She maintained this house for many years, eventually equipping it for science lessons for children.[1]

With several other women, Morgan founded the Summer School Club at Woods Hole in 1913, which is now the Children's School of Science, and served as its first educational chairperson and Science Committee Chair in 1914. She preferred working outdoors with children to conduct experiments and discuss problems.[1]

After her children were old enough, Morgan returned to the laboratory to study Drosophila genetics. Her husband, T.H. Morgan, encouraged her but did not collaborate with her. Instead, he gave her working space in his laboratory, called the "Fly Room," at Columbia University, where she maintained her own Drosophila stocks but held no official position.[1] Her husband and the other male scientists never became comfortable with her presence in the lab, whose atmosphere was "a little like that of an exclusive men's club."[1] Morgan may also have felt isolated because she was older than the other women and was neither outgoing nor talkative, according to Alfred Sturtevant. Because she didn't hold an official position, she never attended a scientific meeting and never presented a paper at a conference.[1]

Morgan and her family moved to California in 1928, where she continued her Drosophila research at the California Institute of Technology in Pasadena while her husband Thomas Hunt Morgan became the division head. Her husband died in 1945; one year afterwards, Morgan received her first official appointment as a research associate at the age of 76.[1] She died in 1952 at the age of 82 in Los Angeles.

• Sampson, L. V. 1894. Die Muskulatur von Chiton. Jenaischen Zeitschrift für Naturwissenschaft 28: 460–468.
• Sampson, L. V. 1895. The musculature of chiton. Journal of Morphology 11:595-628.
• Sampson, L. V. 1900. Unusual modes of breeding and development among anura. Amer. Naturalist 34:687-715.
• Sampson, L. V. 1904. A contribution to the embryology of Hylodes martinicensis. Araer. J. Anat. 3: 473–504.
• Morgan, L. V. 1905. Incomplete anterior regeneration in the absence of the brain in Leploplana litloralis. Biol. Bull. 9:187-193.
• Morgan, L. V. 1906. Regeneration of grafted pieces of planarians. J. Exp. Zool. 3:269-294.
• Morgan, L. V. 1922. Non-criss-cross inheritance in Drosophila melanogaster. Biol. Bull. 42:267-274.
• Morgan, L. V. 1925. Polyploidy in Drosophila melanogaster with two attached X chromosomes" Genetics 10:148-178.
• Morgan, L. V. 1926. Correlation between shape and behavior of a chromosome" Proc. Natl. Acad. Sci 12:180-181.
• Morgan, L. V. 1929. Composites of Drosophila melanogaster. Carnegie Inst. of Wash. Publ. No. 399: 225–296.
• Morgan, L. V. 1931. Proof that bar changes to notbar by unequal crossing-over" Proc. Natl. Acad. Sci 17:270-272.
• Morgan, L. V. 1933. A closed X chromosome in Drosophila melanogaster" Genetics 18:250-283.
• Morgan, L. V. 1938a. Origin of attached-X chromosomes in Drosophila melanogaster and the occurrence of non-disjunction of X's in the male. Amer. Naturalist 72:434-446.
• Morgan, L. V. 1938b. Effects of a compound duplication of the X chromosome of Drosophila melanogaster" Genetics 23:423-462.
• Morgan, L. V. 1939. A spontaneous somatic exchange between non-homologous chromosomes in Drosophila melanogaster" Genetics 24:747-752.
• Morgan, L. V. 1947. A variable phenotype associated with the fourth chromosome of Drosophila melanogaster and affected by heterochromatin" Genetics 32:200-219.
• Morgan, T. H., H. Redfield, and L. V. Morgan. 1943. Maintenance of a Drosophila stock center, in connection with investigations on the germinal material in relation to heredity. Carnegie Inst. Wash. Yearbk. 42:171-174.
• Morgan, T. H., A. H. Sturtevant, and L. V. Morgan. 1945. Maintenance of a Drosophila stock center, in connection with investigations on the germinal material in relation to heredity. Carnegie Inst. Wash. Yearbk. 44:157-160.

• Children's School of Science in Woods Hole, MA.