Praemegaceros

Known from the late Villafranchian of Europe (including Central Italy, Central Romania and Moldova) and the Near East, extending from the Cromer Forest Bed in England to Ubeidiya, Israel, Dmanisi, Georgia and the Azov region, Russia. Croitor suggests an origin in South Asia for this species, descended from taxa possibly referrable to Panolia sp. from the Siwaliks.[7]

Named in 1882 from remains from the Middle Pleistocene Cromer Forest Bed in Norfolk by Edwin Tulley Newton which were originally described in 1872 as belonging to P. verticornis by William Boyd Dawkins. The size is estimated to be around 220 kg, with a mesodont dentition. Croitor suggests that because the pedicles (base of the antlers) are robust and similar to those of giant deer, that P. dawkinsi represents a dwarfed form.[7]

Named by Wolfgang Soergel in 1927 for the species found in the lowest level of the Middle Pleistocene Mosbach locality in Germany.[8] It was included in the “verticornis” group of Azzaroli's 1953 classification. It has been suggested to a be synonym of P. verticornis. However, Croitor suggests that these similarities are the result of parallel evolution, and proposes that P. mosbachensis represents an intermediate form between P. obscurus and P. dawkinsi.[7]

Endemic to Sardinia from the late Middle Pleistocene to the Early Holocene, the ancestor of P. cazioti appears to have dispersed to Sardinia during the Middle Pleistocene, with the earliest well dated records of Praemegaceros at Su Fossu de Cannas in Sardinia being over 450,000 years in age. Other early remains of the genus in Sardinia are referred to the species P. sardus/sardous. The oldest remains assigned to P. cazioti date to approximately 300,000 years ago.[9] P. cazioti is smaller than P. sardus, being slightly larger than a fallow deer. The cranial morphology appears to be unaffected by the insular dwarfism. Two chronologically separated subspecies are known, which are suggested to be chronospecies. P. cazioti cazioti dates to the late Middle Pleistocene and Late Pleistocene and is characterised by smaller brachyodont teeth and P. cazioti algarensis from the end of the Late Pleistocene is larger than P. cazioti cazioti and is characterised by large mesodont upper cheek teeth, and a long premolar series, which are suggested to be adaptions for a grazing diet.[7]

Named by Robert in 1930 for remains from Soleilhac, an early Middle Pleistocene site located in the Massif Central, France. The neotype specimen consists of a partial left frontal with an attached partial antler. The morphology of the preserved antler strongly resembles that of P. cazioti, which suggests a close relationship. Other known sites are from the early Middle Pleistocene of France, Germany, Italy, South Russia and Moldova. This taxon is suggested to be the largest species of Praemegaceros, with an estimated mass of 420 kg. The taxon appears to have been extinct by the late Middle Pleistocene.[7]

Known from the late Early Pleistocene and Middle Pleistocene of Italy, North Greece, Moldova, South Ukraine and the Azov Sea Region. It was named by De Alessandri in 1903 for remains found in North Italy. Psekupsoceros orientalis is a junior synonym of the taxon.[7]

Named by William Boyd Dawkins in 1872 on the basis of a partial antler from the Cromer Forest Bed. Croitor suggests that Cervus belgrandi is a junior synonym.[7] Known from late Early Pleistocene to Middle Pleistocene localities in England, Spain,[10] France,[11] Germany[12] and Italy.[13]