Sardinian warbler

The Sardinian warbler (Curruca melanocephala) is a common and widespread typical warbler from the Mediterranean region. Like most Curruca species, it has distinct male and female plumages. The adult male has a grey back, whitish underparts, black head, white throat and red eyes. Plumages are somewhat variable even in the same locality, with the intensity of a reddish hue on upper- and/or underside that varies from absent to (in some subspecies) pronounced. The female is mainly brown above and buff below, with a grey head. The Sardinian warbler's song is fast and rattling, and is very characteristic of the Mediterranean areas where this bird breeds.

Sardinian warbler
Adult male, Methana peninsula, Greece
Female, Souss-Massa National Park, Morocco
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Class: Aves
Order: Passeriformes
Family: Sylviidae
Genus: Curruca
Species:
C. melanocephala
Binomial name
Curruca melanocephala
(Gmelin, JF, 1789)
Subspecies

2-6, see text

Synonyms

Sylvia melanocephala

Taxonomy and systematics

The Sardinian warbler was formally described in 1789 by the German naturalist Johann Friedrich Gmelin in his revised and expanded edition of Carl Linnaeus's Systema Naturae. He placed it with the wagtails in the genus Motacilla and coined the binomial name Motacilla melanocephala.[2][3] Gmelin based his entry on a description by the Italian zoologist Francesco Cetti in his book Gli uccelli di Sardegna (The Birds of Sardinia) that was published in 1776. This was the second volume of his Storia naturale di Sardegna. Cetti did not use scientific names for the species.[4] When a translation of Cetti's three volumes were published in German, the translator, David Piesch, and the editor, Nathanael Gottfried Leske, included an Appendix at the end of the third volume (published in 1784) in which they proposed the binomial name Motacilla melanocephala, the identical name that was later adopted by Gmelin.[5][6] In spite of this earlier publication, as of 2023 Gmelin is treated as the authority for the species.[7]

The species was formerly placed in the genus Sylvia that was introduced in 1769 by the Italian naturalist Giovanni Antonio Scopoli.[8] The genus name is from Modern Latin silvia, a woodland sprite, related to silva, a wood. The specific melanocephala is from Ancient Greek melas, "black", and kephale, "head".[9] Currently, the Sardinian warbler is placed in the genus Curruca by the IOC, along with most of the species formerly classified in the genus Sylvia.[10]

Together with Menetries's warbler the Sardinian warbler forms a superspecies. Both have white malar areas and light throats, and otherwise black heads in adult males, as well as a naked ring around the eye. The eastern subalpine warbler, which seems the superspecies' closest relative, has a dark throat and breast and a dark gray upper head in males, but otherwise shares these characters. These three species are related to a dark-throated superspecies consisting of Rüppell's warbler and the Cyprus warbler, which also share the white malar area with blackish above.[11]

This bird may be considered a superspecies, divided into the western Curruca melanocephala and Curruca momus from the more arid regions of the Near East and adjacent Africa.

Subspecies

The geographical variation in the Sardinian warbler conforms to some extent with Gloger's rule, though not as strongly as in some other typical warblers. The validity of leucogastra and norissae is not accepted by some authors, and valverdei has been described very recently. On the other hand, leucogastra might be more than one subspecies.[12]

  • Curruca melanocephala melanocephala (Gmelin, 1789)
Iberia across the northern Mediterranean to western Turkey. Extends into the Maghreb from Iberia, and into Libya from Italy via Sicily. Migrates to the Sahel and oases in the Sahara in winter.[12]
Large, long wings, tail tip rather pointed. A dark form, usually lacking any reddish in males but flanks extensively grey. Females' uppersides vary between deep olive brown and greyish olive.[12]
  • Curruca melanocephala leucogastra (Ledru, 1810) - often included in melanocephala; phylogenetic status requires review[12]
Canary Islands, resident, probably some vagrancy between eastern islands and Maghreb.[12]
Medium size, short-winged and large-billed. Tenerife and La Palma (western) birds are most distinct, being dark above with some rusty/beige hue on the underside in males. Eastern birds (Fuerteventura, Lanzarote and Gran Canaria) are more like melanocephala and momus but differ in measurements.[12]
  • Curruca melanocephala momus (Hemprich & Ehrenberg, 1833)
Near East. Resident, some local movements.[12]
Smallish, short-winged. Varies between brownish grey and rusty above; underside almost always has reddish hue. Females rusty to rusty olive.[12]
  • Curruca melanocephala norissae Nicoll, 1917: Fayyum warbler - probably only a local morph of momus[12]
Nile Delta region. Extinct since around 1940.[12]
Like momus, but tend to be very reddish.[12]
  • Curruca melanocephala valverdei Cabot & Urdiales, 2005 - recently split from melanocephala
From Tiznit (Morocco) south to the Tropic of Cancer, inland to the edge of the Sahara. Resident, but some seasonal movements.[12]
Medium-sized, tail tip quite square. A very pure-colored form, the palest subspecies. Undersides clean white. Matte black cap in males. Juveniles decidedly sandy.[12]

Distribution and habitat

It breeds in the southernmost areas of Europe and just into Asia in Turkey and the eastern end of the Mediterranean. This small passerine bird, unlike most "warblers", is not particularly migratory, but some birds winter in North Africa, and it occurs as a vagrant well away from the breeding range, as far as Great Britain.

Behaviour and ecology
Curruca melanocephala - MHNT
Cuculus canorus (cuckoo) egg in a Sylvia melanocephala clutch MHNT

This is a bird of open country and cultivation, with bushes for nesting. The nest is built in low shrub or brambles, and 3-6 eggs are laid. Like most "warblers", it is insectivorous, but will also take berries and other soft fruit.

References
  1. BirdLife International (2018). "Curruca melanocephala". IUCN Red List of Threatened Species. 2018. doi:10.2305/IUCN.UK.2018-2.RLTS.T22716959A132113832.en. Retrieved 20 October 2022.
  2. Gmelin, Johann Friedrich (1789). Systema naturae per regna tria naturae : secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1, Part 2 (13th ed.). Lipsiae [Leipzig]: Georg. Emanuel. Beer. p. 970.
  3. Mayr, Ernst; Cottrell, G. William, eds. (1986). Check-list of Birds of the World. Vol. 11. Cambridge, Massachusetts: Museum of Comparative Zoology. p. 281.
  4. Cetti, Francesco (1776). Storia naturale di Sardegna (in Italian). Vol. 2: Gli uccelli di Sardegna. Sassari: G. Piattoli. p. 218.
  5. Cetti, Francesco (1784). "Appendix". Naturgeschichte von Sardinien (in German). Vol. 3: Geschichte der Amphibien und Fische. Translated by Piesch, David. Notes by Leske, Nathanael Gottfried. Leipzig: Müller. p. 18.
  6. Mlíkovský, Jiří (2020). "The original description of Sylvia melanocephala (Aves: Sylviidae), with notes on David Piesch's birds of Sardinia (1784)". Rivista italiana di Ornitologia - Research in Ornithology. 90 (2): 95–96. doi:10.4081/rio.2020.489. S2CID 234141771.
  7. Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (January 2023). "Sylviid babblers, parrotbills, white-eyes". IOC World Bird List Version 13.1. International Ornithologists' Union. Retrieved 8 May 2023.
  8. Scopoli, Giovanni Antonio (1769). Annus I historico-naturalis (in Latin). Lipsiae (Leipzig): C.G. Hilscheri. p. 154.
  9. Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 246, 376. ISBN 978-1-4081-2501-4.
  10. Dickinson, E.C.; Christidis, L., eds. (2014). The Howard & Moore Complete Checklist of the Birds of the World, Volume 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. pp. 509–512. ISBN 978-0-9568611-2-2.
  11. Helbig (2001), Jønsson & Fjeldså (2006)
  12. Cabot & Urdiales (2005)
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