Sclerosomatidae

The Sclerosomatidae are a family of harvestmen with about 1,300 known species.[1] One former subfamily has been recently removed to form a new family, Globipedidae.[2]

Sclerosomatidae
Temporal range:
Hadrobunus grandis
Scientific classification
Kingdom:
Animalia
Phylum:
Arthropoda
Class:
Arachnida
Order:
Opiliones
Suborder:
Eupnoi
Superfamily:
Phalangioidea
Family:
Sclerosomatidae

Simon, 1879
Subfamilies

Gagrellinae (117 genera)
Gyantinae (3 genera)
Leiobuninae (13 genera)
Sclerosomatinae (10 genera)

Diversity
4 subfamilies, c. 150 genera

Name

The name is combined from Ancient Greek skleros "hard" and soma "body".[3]

Genera
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  • Abaetetuba Tourinho-Davis, 2004 (5 species)
  • Adungrella Roewer, 1955 (3 species)
  • Aguilaia González-Sponga, 2003 (1 species; Venezuela)
  • Akalpia Roewer, 1915 (2 species; India and Japan)
  • Altobunus Roewer, 1910 (3 species; Celebes and Philippines)
  • Amazonesia Soares, 1970 (2 species)
  • Antigrella Roewer, 1954 (1 species)
  • Aurivilliola Roewer, 1910 (17 species; South and Southeast Asia)
  • Azucarella Roewer, 1959 (1 species)
  • Bakerinulus Roewer, 1955 (1 species)
  • Bastia Roewer, 1910 (3 species; South and Southeast Asia)
  • Bastioides Mello-Leitão, 1931 (1 species; South America)
  • Baturitia Roewer, 1931 (1 species; Sunda Islands)
  • Bellonia González-Sponga, 2003 (1 species; Venezuela)
  • Biceropsis Roewer, 1935 (1 species; Burma)
  • Bonthainia Roewer, 1913 (5 species; South and Southeast Asia)
  • Bullobunus Roewer, 1910 (7 species; Philippines)
  • Caluga Roewer, 1959 (1 species)
  • Cardenalia Kury, 2017 (1 species; Venezuela)
  • Carinobius Roewer, 1955 (1 species)
  • Carmenia Roewer, 1915 (1 species; Colombia)
  • Carmichaelus Roewer, 1929 (1 species; India)
  • Ceratobunellus Roewer, 1911 (3 species; India)
  • Ceratobunoides Roewer, 1923 (2 species; Sumatra)
  • Ceratobunus Thorell, 1889 (13 species)
  • Cervibunus Roewer, 1912 (3 Species; South and Southeast Asia)
  • Chasenella Roewer, 1955 (2 species; Borneo)
  • Chebabius Roewer, 1935 (1 species; Burma)
  • Coonoora Roewer, 1929 (1 species; India)
  • Dentobunus Roewer, 1910 (33 species; Southeast Asia)
  • Diangathia Roewer, 1955 (1 species)
  • Echinobunus Roewer, 1912 (1 species; Java)
  • Euceratobunus Roewer, 1923 (1 species; India)
  • Eugagrella Roewer, 1910 (25 species; South and Southeast Asia)
  • Euzaleptus Roewer, 1911 (4 species; South and Southeast Asia)
  • Fesa Roewer, 1953 (1 species)
  • Gagrella Stoliczka, 1869 (226 species; Asia)
  • Gagrellenna Roewer, 1929 (1 species; India)
  • Gagrellina Roewer, 1913 (1 species; Celebes)
  • Gagrellissa Roewer, 1931 (1 species; Sunda Islands)
  • Gagrellopsis Sato & Suzuki, 1939 (1 species; Japan)
  • Gagrellula Roewer, 1910 (67 species; Asia)
  • Garleppa Roewer, 1912 (4 species; South America)
  • Geaya Roewer, 1910 (82 species; Latin America)
  • Globulosoma Martens, 1987 (2 species; Nepal)
  • Guaranobunus Ringuelet, 1954 (1 species; Argentina)
  • Hamitergum Crawford, 1992 (1 species; Russia)
  • Harmanda Roewer, 1910 (15 species; Asia)
  • Harmandina Schenkel, 1954 (1 species)
  • Hehoa Roewer, 1929 (1 species; India)
  • Heterogagrella Roewer, 1954 (2 species; Southeast Asia)
  • Hexazaleptus Suzuki, 1966 (1 species)
  • Hexomma Thorell, 1876 (9 species; Southeast Asia)
  • Himaldroma Martens, 1987 (2 species; Nepal)
  • Himalzaleptus Martens, 1987 (1 species; Nepal)
  • Holcobunus Roewer, 1910 (26 or 27 species; Latin America)
  • Holmbergiana Mello-Leitão, 1931 (3 species; South America)
  • Hologagrella Roewer, 1910 (7 species; Southeast Asia)
  • Hypogrella Roewer, 1955 (1 species)
  • Hypsibunus Thorell, 1891 (11 species; India & Southeast Asia)
  • Jussara Mello-Leitão, 1935 (10 species; South America)
  • Koyamaia Suzuki, 1972 (1 species)
  • Krusa Goodnight & Goodnight, 1947 (10 species; Mexico)
  • Krusella Roewer, 1953 (1 species; Venezuela)
  • Liopagus Chamberlin, 1916 (1 species)
  • Marthana Thorell, 1891 (24 species; Southeast Asia)
  • Melanopa Thorell, 1889 (30 species; India, Southeast Asia)
  • Melanopella Roewer, 1931 (3 species; Sunda Islands)
  • Melanopula Roewer, 1929 (5 species; South and Southeast Asia)
  • Metadentobunus Roewer, 1915 (3 species; Taiwan)[4]
  • Metahehoa Suzuki, 1985 (1 species)
  • Metasyleus Roewer, 1955 (3 species)
  • Metaverpulus Roewer, 1912 (7 species; South Asia)
  • Metazaleptus Roewer, 1912 (8 species; South and Southeast Asia)
  • Microzaleptus Roewer, 1955 (1 species)
  • Munequita Mello-Leitão, 1941 (1 species; Brazil)
  • Neogagrella Roewer, 1913 (3 species; South and Southeast Asia)
  • Nepalgrella Martens, 1987 (2 species; Nepal)
  • Nepalkanchia Martens, 1990 (2 species; Nepal)
  • Obigrella Roewer, 1955 (1 species)
  • Octozaleptus Suzuki, 1966 (1 species)
  • Onostemma Mello-Leitão, 1938 (2 species; Brazil)
  • Oobunus Kishida, 1930 (1 species)
  • Orissula Roewer, 1955 (1 species)
  • Padangrella Roewer, 1954 (1 species)
  • Palniella Roewer, 1929 (1 species; India)
  • Paradentobunus Roewer, 1915 (1 species; India)
  • Paragagrella Roewer, 1912 (5 species; South and Southeast Asia)
  • Paragagrellina Schenkel, 1963 (1 species)
  • Parageaya Mello-Leitão, 1933 (6 species; Latin America)
  • Paratamboicus Mello-Leitão, 1940 (30 species; South America)
  • Paraumbogrella Suzuki, 1963 (1 species)
  • Paruleptes Soares, 1970 (1 species)
  • Pectenobunus Roewer, 1910 (4 species; South America)
  • Pergagrella Roewer, 1954 (1 species)
  • Pokhara Suzuki, 1970 (7 species; Nepal)
  • Prionostemma Pocock, 1903 (114 species; Mexico, Central America, and South America)
  • Prodentobunus Roewer, 1923 (4 species; Asia)
  • Psammogeaya Mello-Leitão, 1946 (1 species; Uruguay)
  • Psathyropus L. Koch, 1878 (40 species; Asia)
  • Pseudarthromerus Karsch, 1892 (1 species)
  • Pseudogagrella Redikortsev, 1936 (13 species; East Asia)
  • Pseudomelanopa Suzuki, 1974 (2 species; East Asia)
  • Pseudosystenocentrus Suzuki, 1985 (1 species)
  • Romerella Goodnight & Goodnight, 1943 (5 species; Mexico and South America)
  • Sarasinia Roewer, 1913 (1 species; Celebes)
  • Sataria Roewer, 1915 (3 species; India)
  • Scotomenia Thorell, 1889 (1 species; Burma)
  • Sericicorpus Martens, 1987 (1 species; Nepal)
  • Sinadroma Roewer, 1955 (1 species)
  • Syleus Thorell, 1876 (2 species; India)
  • Syngagrella Roewer, 1913 (1 species; Celebes)
  • Systenocentrus Simon, 1886 (6 species; Southeast and East Asia)
  • Tamboicus Roewer, 1912 (3 species; South America)
  • Taperina Roewer, 1953 (2 species)
  • Tetraceratobunus Roewer, 1915 (3 species; South Asia)
  • Thunbergia Martens, 2020 (4 species; South America)
  • Toragrella Roewer, 1955 (2 species)
  • Trachyrhinus Weed, 1892 (7 species; North America: Mexico and United States)
  • Umbogrella Roewer, 1955 (1 species)
  • Umbopilio Roewer, 1956 (2 species)
  • Vaho González-Sponga, 2003 (1 species)
  • Valle González-Sponga, 2003 (3 species)
  • Varinodulia Canals, 1935 (1 species)
  • Verpulus Simon, 1901 (13 species; South and Southeast Asia)
  • Verrucobunus Roewer, 1931 (3 species; East and Southeast Asia)
  • Xerogrella Martens, 1987 (1 species; Nepal)
  • Zaleptiolus Roewer, 1955 (4 species; Asia)
  • Zaleptulus Roewer, 1955 (3 species)
  • Zaleptus Thorell, 1876 (72 species; Asia and Australia)
  • Gyantinae Silhavý, 1946
  • Gyas Simon, 1879 (1 species; Alps)
  • Gyoides Martens, 1982 (6 species; Nepal)
  • Rongsharia Roewer, 1957 (3 species; Nepal)
  • Leiobuninae Banks, 1893
  • †Amauropilio Mello-Leitão, 1937 (fossil) (2 species; fossils found in Colorado)
  • Cosmobunus Simon, 1879 (5 species; Spain and Mexico)
  • Dilophiocara Redikorzev, 1931 (2 species)
  • Eumesosoma Cokendolpher, 1980 (6 species; United States)
  • Eusclera Roewer, 1910 (2 species; China and India)
  • Hadrobunus Banks, 1900 (2 species; United States)
  • Leiobunum C.L.Koch, 1839 (126 species - 2 extinct fossils; worldwide)
  • Leuronychus Banks, 1900 (2 species; North America)
  • Microliobunum Roewer, 1912 (2 species; Middle East)
  • Micronelima Schenkel, 1938 (1 species; Spain)
  • Nelima Roewer, 1910 (44 species; worldwide)
  • Paranelima Caporiacco, 1938 (6 species; Mexico and Central America)
  • Schenkeliobunum Starega, 1964 (1 species)
  • Togwoteeus Roewer, 1952 (1 species; United States)
  • Sclerosomatinae Simon, 1879

Removed
  • Caiza Roewer, 1925. Subjective synonym of Pectenobunus Roewer, 1910) in Tourinho, 2004a

References
  1. Sclerosomatidae Species List at Joel Hallan's Biology Catalog. Texas A&M University, archived from the original on 13 October 2014, retrieved 30 August 2015
  2. Kury, A.B. & J.C. Cokendolpher (2020) Chapter 9. A new family from the New World (Eupnoi: Phalangioidea). In: WCO-Lite: online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0 — Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018 (Kury et al., ed.): 52-54.
  3. Tourinho, Ana Lúcia (2007): Sclerosomatidae. Simon, 1879. In: Pinto-da-Rocha et al. 2007: 127ff
  4. Chen, S. L.; Shih, H. T. (2018). "A review of the harvestman genus Metadentobunus (Opiliones: Sclerosomatidae: Gagrellinae) with a description of a new species from Taiwan". Zoological Studies. 57.
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