Phlegmariurus

Phlegmariurus is a genus of lycophyte plants in the family Lycopodiaceae.[1] The genus is recognized in the Pteridophyte Phylogeny Group classification of 2016 (PPG I),[2] but not by some other sources, which keep it in a broadly defined Huperzia.

Phlegmariurus
Phlegmariurus phlegmaria growing in the Cambridge University Botanic Garden
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Lycophytes
Class: Lycopodiopsida
Order: Lycopodiales
Family: Lycopodiaceae
Subfamily: Huperzioideae
Genus: Phlegmariurus
(Herter) Holub[1]
Species

See text.

Taxonomy

The genus was first described in 1909 by Wilhelm Herter as the section Phlegmariurus of the genus Lycopodium. The section was elevated to a genus by Josef Ludwig Holub in 1964.[3]

Within the family Lycopodiaceae, Phlegmariurus is placed in the subfamily Huperzioideae. A phylogenetic study in 2016, employing both molecular and morphological data, concluded that either a one-genus or a three-genus division of the subfamily produced monophyletic taxa. The authors preferred the three-genus division, recognizing Huperzia, Phlegmariurus and Phylloglossum. Their preferred hypothesis for the relationships of the three genera was:[4]

Huperzioideae

Phylloglossum

Huperzia

Phlegmariurus

The majority of the species formerly placed in a broadly defined Huperzia belong in Phlegmariurus.[4] However, the genera are difficult to separate morphologically, and others have preferred the one-genus division of the subfamily.[1]

Species

As of January 2023, the Checklist of Ferns and Lycophytes of the World accepted over 300 species:[1]

  • Phlegmariurus acerosus (Sw.) B.Øllg.
  • Phlegmariurus acifolius (Rolleri) B.Øllg.
  • Phlegmariurus acutus (Rolleri) B.Øllg.
  • Phlegmariurus affinis (Trevis.) B.Øllg.
  • Phlegmariurus afromontanus Pic. Serm.
  • Phlegmariurus ambrensis (Rakotondr.) A.R.Field & Bauret
  • Phlegmariurus amentaceus (B.Øllg.) B.Øllg.
  • Phlegmariurus andinus (Rosenst.) B.Øllg.
  • Phlegmariurus aqualupianus (Spring) B.Øllg.
  • Phlegmariurus arcuatus (B.Øllg.) B.Øllg.
  • Phlegmariurus ascendens (Nessel) B.Øllg.
  • Phlegmariurus attenuatus (Spring) B.Øllg.
  • Phlegmariurus australis (Willd.) A.R.Field
  • Phlegmariurus austroecuadoricus (B.Øllg.) B.Øllg.
  • Phlegmariurus austrosinicus (Ching) Li Bing Zhang
  • Phlegmariurus axillaris (Roxb.) comb. ined., currently Huperzia axillaris (Roxb.) Rothm.
  • Phlegmariurus badinianus (B.Øllg. & P.G.Windisch) B.Øllg.
  • Phlegmariurus balansae (Herter) A.R.Field & Bostock
  • Phlegmariurus bampsianus (Pic. Serm.) A.R.Field & Bostock
  • Phlegmariurus banayanicus (Herter) A.R.Field & Bostock
  • Phlegmariurus beitelii (B.Øllg.) B.Øllg.
  • Phlegmariurus biformis (Hook.) B.Øllg.
  • Phlegmariurus billardierei (Spring) Trevis.
  • Phlegmariurus binervius (Herter) B.Øllg.
  • Phlegmariurus bolanicus (Rosenst.) A.R.Field & Bostock
  • Phlegmariurus brachiatus (Maxon) B.Øllg.
  • Phlegmariurus brachystachys (Baker) A.R.Field & Bostock
  • Phlegmariurus bradeorum (Christ) B.Øllg.
  • Phlegmariurus brevifolius (Hook. & Grev.) B.Øllg.
  • Phlegmariurus brongniartii (Spring) B.Øllg.
  • Phlegmariurus buesii (Herter) B.Øllg.
  • Phlegmariurus callitrichifolius (Mett.) B.Øllg.
  • Phlegmariurus campianus (B.Øllg.) B.Øllg.
  • Phlegmariurus cancellatus (Spring) Ching
  • Phlegmariurus capellae (Herter) B.Øllg.
  • Phlegmariurus capillaris (Sodiro) B.Øllg.
  • Phlegmariurus carinatus (Poir.) Ching
  • Phlegmariurus catacachiensis (Nessel) B.Øllg.
  • Phlegmariurus cavifolius (C.Chr.) A.R.Field & Bostock
  • Phlegmariurus chamaeleon (Herter) B.Øllg.
  • Phlegmariurus changii T.Y.Hsieh
  • Phlegmariurus chiricanus (Maxon) B.Øllg.
  • Phlegmariurus christii (Silveira) B.Øllg.
  • Phlegmariurus ciliatospiculatus B.Øllg.
  • Phlegmariurus ciliolatus (B.Øllg.) B.Øllg.
  • Phlegmariurus cleefianus (B.Øllg.) B.Øllg.
  • Phlegmariurus cocouyensis B.Øllg.
  • Phlegmariurus colanensis (B.Øllg.) B.Øllg.
  • Phlegmariurus columnaris (B.Øllg.) B.Øllg.
  • Phlegmariurus comans (Herter ex Nessel) B.Øllg.
  • Phlegmariurus compactus (Hook.) B.Øllg.
  • Phlegmariurus copelandianus (R.C.Y.Chou & Bartlett) A.R.Field
  • Phlegmariurus coralium (Spring) A.R.Field & Bostock
  • Phlegmariurus costaricensis (Herter) comb. ined., currently Huperzia costaricensis (Herter) Holub
  • Phlegmariurus crassus (Humb. & Bonpl. ex Willd.) B.Øllg.
  • Phlegmariurus creber (Alderw.) A.R.Field & Bostock
  • Phlegmariurus crucis-australis (Herter) B.Øllg.
  • Phlegmariurus cruentus (Spring) B.Øllg.
  • Phlegmariurus cryptomerinus (Maxim.) Satou
  • Phlegmariurus cuernavacensis (Underw. & F.E.Lloyd) B.Øllg.
  • Phlegmariurus cumingii (Nessel) B.Øllg.
  • Phlegmariurus cuneifolius (Hieron.) B.Øllg.
  • Phlegmariurus cunninghamioides (Hayata) Ching
  • Phlegmariurus curiosus (Herter) A.R.Field & Bostock
  • Phlegmariurus curvifolius (Kunze) B.Øllg.
  • Phlegmariurus dacrydioides (Baker) A.R.Field & Bostock
  • Phlegmariurus dalhousieanus (Spring) A.R.Field & Bostock
  • Phlegmariurus darwinianus (Herter ex Nessel) B.Øllg.
  • Phlegmariurus delbrueckii (Herter) A.R.Field & Bostock
  • Phlegmariurus deminuens (Herter) B.Øllg.
  • Phlegmariurus dentatus (Herter) Arana
  • Phlegmariurus dianae (Herter) B.Øllg.
  • Phlegmariurus dichaeoides (Maxon) B.Øllg.
  • Phlegmariurus dichotomus (Jacq.) W.H.Wagner
  • Phlegmariurus dielsii (Herter) A.R.Field & Bostock
  • Phlegmariurus divergens (Alderw.) A.R.Field & Testo
  • Phlegmariurus durus (Pic. Serm.) A.R.Field & Bostock
  • Phlegmariurus echinatus (Spring) B.Øllg.
  • Phlegmariurus ellenbeckii (Nessel) A.R.Field & Bostock
  • Phlegmariurus elmeri (Herter) A.R.Field & Bostock
  • Phlegmariurus engleri (Hieron. & Herter) B.Øllg.
  • Phlegmariurus eremorum (Rolleri) B.Øllg.
  • Phlegmariurus ericifolius (C.Presl) B.Øllg.
  • Phlegmariurus erythrocaulos (Fée) B.Øllg.
  • Phlegmariurus espinosanus (B.Øllg.) B.Øllg.
  • Phlegmariurus eversus (Poir.) B.Øllg.
  • Phlegmariurus fargesii (Herter) Ching
  • Phlegmariurus filicaulos (Copel.) A.R.Field & Testo
  • Phlegmariurus filiformis (Sw.) W.H.Wagner
  • Phlegmariurus firmus (Mett.) B.Øllg.
  • Phlegmariurus flagellaceus (Kuhn) A.R.Field & Bostock
  • Phlegmariurus flexibilis (Fée) B.Øllg.
  • Phlegmariurus foliaceus (Maxon) B.Øllg.
  • Phlegmariurus foliosus (Copel.) A.R.Field & Bostock
  • Phlegmariurus fontinaloides (Spring) B.Øllg.
  • Phlegmariurus fordii (Baker) Ching
  • Phlegmariurus friburgensis (Herter ex Nessel & Hoehne) B.Øllg.
  • Phlegmariurus funiculus (Herter) comb. ined., currently Huperzia funicula (Herter) J.P.Roux
  • Phlegmariurus funiformis (Cham. ex Spring) B.Øllg.
  • Phlegmariurus gagnepainianus (Herter) A.R.Field & Bostock
  • Phlegmariurus galapagensis (O.J.Hamann) B.Øllg.
  • Phlegmariurus giganteus (Herter) A.R.Field & Bostock
  • Phlegmariurus gnidioides (L. fil.) A.R.Field & Bostock
  • Phlegmariurus goebelii (Nessel) A.R.Field & Bostock
  • Phlegmariurus gracilis A.Rojas & R.Calderón
  • Phlegmariurus guandongensis Ching
  • Phlegmariurus gunturensis (Alderw.) A.R.Field & Bostock
  • Phlegmariurus haeckelii (Herter) A.R.Field & Testo
  • Phlegmariurus haitensis (Herter) comb. ined., currently Huperzia haitensis (Herter) Holub
  • Phlegmariurus hamiltonii (Spreng.) Á.Löve & D.Löve
  • Phlegmariurus harmsii (Nessel) A.R.Field & Bostock
  • Phlegmariurus hartwegianus (Spring) B.Øllg.
  • Phlegmariurus hastatus (B.Øllg.) B.Øllg.
  • Phlegmariurus hellwigii (Warb.) A.R.Field & Bostock
  • Phlegmariurus hemleri (Nessel) B.Øllg.
  • Phlegmariurus henryi (Baker) Ching
  • Phlegmariurus heterocarpos (Fée) B.Øllg.
  • Phlegmariurus heteroclitus (Desv. ex Poir.) B.Øllg.
  • Phlegmariurus hexastichus (B.Øllg. & P.G.Windisch) B.Øllg.
  • Phlegmariurus hildebrandtii (Herter) A.R.Field & Bauret
  • Phlegmariurus hippurideus (Christ) B.Øllg.
  • Phlegmariurus hippuris (Desv. ex Poir.) A.R.Field & Testo
  • Phlegmariurus hoffmannii (Maxon) B.Øllg.
  • Phlegmariurus hohenackeri (Herter) B.Øllg.
  • Phlegmariurus holstii (Hieron.) A.R.Field & Bostock
  • Phlegmariurus homocarpus (Herter) B.Øllg.
  • Phlegmariurus horizontalis (Nessel) A.R.Field & Bostock
  • Phlegmariurus huberi (B.Øllg.) B.Øllg.
  • Phlegmariurus hugoi B.Øllg.
  • Phlegmariurus humbertii (Nessel) A.R.Field & Bostock
  • Phlegmariurus humbertii-henrici (Herter) A.R.Field & Bostock
  • Phlegmariurus hypogaeus (B.Øllg.) B.Øllg.
  • Phlegmariurus hystrix (Herter) B.Øllg.
  • Phlegmariurus idroboi B.Øllg.
  • Phlegmariurus iminii Kiew
  • Phlegmariurus insularis (Carmich.) comb. ined., currently Huperzia insularis (Carmich.) Rothm.
  • Phlegmariurus intermedius (Trevis.) B.Øllg.
  • Phlegmariurus itambensis (B.Øllg. & P.G.Windisch) B.Øllg.
  • Phlegmariurus jaegeri (Herter) A.R.Field & Bostock
  • Phlegmariurus josesantae B.Øllg.
  • Phlegmariurus kajewskii (Copel.) A.R.Field & Testo
  • Phlegmariurus killipii (Herter) B.Øllg.
  • Phlegmariurus kuesteri (Nessel) B.Øllg.
  • Phlegmariurus lancifolius (Maxon) B.Øllg.
  • Phlegmariurus lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock
  • Phlegmariurus lecomteanus (Nessel) A.R.Field & Bostock
  • Phlegmariurus ledermannii (Herter) A.R.Field & Bostock
  • Phlegmariurus lehnertii Testo
  • Phlegmariurus lignosus (Herter) B.Øllg.
  • Phlegmariurus lindenii (Spring) B.Øllg.
  • Phlegmariurus linifolius (L.) B.Øllg.
  • Phlegmariurus llanganatensis (B.Øllg.) B.Øllg.
  • Phlegmariurus lockyeri (D.Jones & B.Gray) A.R.Field & Bostock
  • Phlegmariurus loefgrenianus (Silveira) B.Øllg.
  • Phlegmariurus longus (Copel.) A.R.Field & Bostock
  • Phlegmariurus loxensis (B.Øllg.) B.Øllg.
  • Phlegmariurus luteynii B.Øllg.
  • Phlegmariurus macbridei (Herter) B.Øllg.
  • Phlegmariurus macgregorii (Baker) A.R.Field & Bostock
  • Phlegmariurus macrostachys (Hook. ex Spring) N.C.Nair & S.R.Ghosh
  • Phlegmariurus magnusianus (Herter) A.R.Field & Testo
  • Phlegmariurus mandiocanus (Raddi) B.Øllg.
  • Phlegmariurus mannii (Hillebr.) W.H.Wagner
  • Phlegmariurus marsupiiformis (D.Jones & B.Gray) A.R.Field & Bostock
  • Phlegmariurus martii (Wawra) B.Øllg.
  • Phlegmariurus megastachyus (Baker) A.R.Field & Bostock
  • Phlegmariurus melanesicus (Brownlie) A.R.Field & Testo
  • Phlegmariurus merrillii (Herter) A.R.Field & Bostock
  • Phlegmariurus mesoamericanus (B.Øllg.) B.Øllg.
  • Phlegmariurus mexicanus (Herter) B.Øllg.
  • Phlegmariurus mildbraedii (Herter) A.R.Field & Bostock
  • Phlegmariurus mingcheensis Ching
  • Phlegmariurus minutifolius (Alderw.) A.R.Field & Bostock
  • Phlegmariurus mirabilis (Willd.) A.R.Field & Testo
  • Phlegmariurus mollicomus (Spring) B.Øllg.
  • Phlegmariurus monticola Kiew
  • Phlegmariurus mooreanus (Sander ex Baker) B.Øllg.
  • Phlegmariurus multifarius (Alderw.) A.R.Field & Bostock
  • Phlegmariurus myrsinites (Lam.) B.Øllg.
  • Phlegmariurus myrtifolius (G.Forst.) A.R.Field & Bostock
  • Phlegmariurus myrtuosus (Spring) B.Øllg.
  • Phlegmariurus neocaledonicus (Nessel) A.R.Field & Bostock
  • Phlegmariurus nesselii (Brause ex Nessel) B.Øllg.
  • Phlegmariurus niligaricus (Spring) A.R.Field & Bostock
  • Phlegmariurus nudus (Christ ex Nessel & Hoehne) B.Øllg.
  • Phlegmariurus nummularifolius (Blume) Ching
  • Phlegmariurus nutans (Brack.) W.H.Wagner
  • Phlegmariurus nyalamensis (Ching & S.K.Wu) H.S.Kung & L.B.Zhang
  • Phlegmariurus obovalifolius (Bonap.) A.R.Field & Testo
  • Phlegmariurus obtusifolius (P.Beauv.) A.R.Field & Bostock
  • Phlegmariurus ocananus (Herter) B.Øllg.
  • Phlegmariurus oceanianus (Herter) A.R.Field & Bostock
  • Phlegmariurus oellgaardii (A.Rojas) B.Øllg.
  • Phlegmariurus oltmannsii (Herter ex Nessel) A.R.Field & Bostock
  • Phlegmariurus ophioglossoides (Lam.) A.R.Field & Bostock
  • Phlegmariurus orizabae (Underw. & F.E.Lloyd) B.Øllg.
  • Phlegmariurus ovatifolius (Ching) W.M.Chu ex H.S.Kung & Li Bing Zhang
  • Phlegmariurus pachyphyllus (Kuhn ex Herter) A.R.Field & Testo
  • Phlegmariurus pachyskelos B.Øllg.
  • Phlegmariurus parksii (Copel.) A.R.Field & Bostock
  • Phlegmariurus patentissimus (Alderw.) A.R.Field & Bostock
  • Phlegmariurus pecten (Baker) A.R.Field & Bostock
  • Phlegmariurus perrierianus (Tardieu) A.R.Field & Bostock
  • Phlegmariurus pflanzii (Nessel) B.Øllg.
  • Phlegmariurus phlegmaria (L.) Holub
  • Phlegmariurus phlegmarioides (Gaudich.) A.R.Field & Bostock
  • Phlegmariurus phylicifolius (Desv. ex Poir.) B.Øllg.
  • Phlegmariurus phyllanthus (Hook. & Arn.) R.D.Dixit
  • Phlegmariurus picardae (Krug) B.Øllg.
  • Phlegmariurus pichianus (Tardieu) A.R.Field & Bostock
  • Phlegmariurus pinifolius (Trevis.) Kiew
  • Phlegmariurus pithyoides (Schltdl. & Cham.) B.Øllg.
  • Phlegmariurus pittieri (Christ) B.Øllg.
  • Phlegmariurus podocarpensis B.Øllg.
  • Phlegmariurus polycarpos (Kunze) B.Øllg.
  • Phlegmariurus polycladus (Sodiro) comb. ined., currently Huperzia polyclada (Sodiro) Rolleri & Deferrari
  • Phlegmariurus polydactylus (B.Øllg.) B.Øllg.
  • Phlegmariurus polylepidetorum (B.Øllg.) B.Øllg.
  • Phlegmariurus portoricensis (Underw. & F.E.Lloyd) comb. ined., currently Lycopodium portoricense Underw. & F.E.Lloyd
  • Phlegmariurus pringlei (Underw. & F.E.Lloyd) B.Øllg.
  • Phlegmariurus proliferus (Blume) A.R.Field & Bostock
  • Phlegmariurus pruinosus (Hieron. & Herter) B.Øllg.
  • Phlegmariurus pseudovarius (Brownlie) Rouhan & A.R.Field
  • Phlegmariurus pulcherrimus (Hook. & Grev.) Á. & D.Löve
  • Phlegmariurus pungentifolius (Silveira) B.Øllg.
  • Phlegmariurus quadrifariatus (Bory) B.Øllg.
  • Phlegmariurus recurvifolius (Rolleri) B.Øllg.
  • Phlegmariurus reflexus (Lam.) B.Øllg.
  • Phlegmariurus regnellii (Maxon) B.Øllg.
  • Phlegmariurus ribourtii (Herter) A.R.Field & Bostock
  • Phlegmariurus riobambensis (Nessel) B.Øllg.
  • Phlegmariurus robustus (Klotzsch) B.Øllg.
  • Phlegmariurus rosenstockianus (Herter) B.Øllg.
  • Phlegmariurus rostrifolius (Silveira) B.Øllg.
  • Phlegmariurus ruber (Cham. & Schltdl.) B.Øllg.
  • Phlegmariurus rubricus (Herter) A.R.Field & Bostock
  • Phlegmariurus rufescens (Hook.) B.Øllg.
  • Phlegmariurus rupicola (Alderw.) A.R.Field & Bostock
  • Phlegmariurus sagasteguianus (B.Øllg.) B.Øllg.
  • Phlegmariurus salvinioides (Herter) Ching
  • Phlegmariurus samoanus (Herter ex Nessel) A.R.Field & Bostock
  • Phlegmariurus sarmentosus (Spring) B.Øllg.
  • Phlegmariurus saururus (Lam.) B.Øllg.
  • Phlegmariurus scabridus (B.Øllg.) B.Øllg.
  • Phlegmariurus schlechteri (E.Pritz.) A.R.Field & Bostock
  • Phlegmariurus schlimii (Herter) B.Øllg.
  • Phlegmariurus schmidtchenii (Hieron.) B.Øllg.
  • Phlegmariurus sellifolius (B.Øllg.) B.Øllg.
  • Phlegmariurus sellowianus (Herter) B.Øllg.
  • Phlegmariurus serpentiformis (Herter) B.Øllg.
  • Phlegmariurus setifolius (Alderw.) A.R.Field & Bostock
  • Phlegmariurus shangsiensis C.Y.Yang
  • Phlegmariurus shingianus R.H.Jiang & X.C.Zhang
  • Phlegmariurus sieberianus (Spring) B.Øllg.
  • Phlegmariurus sieboldii (Miq.) Ching
  • Phlegmariurus silveirae (Herter ex Nessel & Hoehne) B.Øllg.
  • Phlegmariurus silverstonei B.Øllg.
  • Phlegmariurus sintenisii (Herter) B.Øllg.
  • Phlegmariurus societensis (J.W.Moore) A.R.Field
  • Phlegmariurus sooianus Lawalrée
  • Phlegmariurus sotae (Rolleri) B.Øllg.
  • Phlegmariurus sphagnicola B.Øllg.
  • Phlegmariurus squarrosus (G.Forst.) Á. & D.Löve
  • Phlegmariurus staudtii (Nessel) A.R.Field & Bostock
  • Phlegmariurus stemmermanniae A.C.Medeiros & Wagner
  • Phlegmariurus stephani B.Øllg.
  • Phlegmariurus strictus (Baker) A.R.Field & Bostock
  • Phlegmariurus subfalciformis (Alderw.) A.R.Field & Bostock
  • Phlegmariurus subtubulosus (Herter) comb. ined., currently Huperzia subtubulosa (Herter) Holub
  • Phlegmariurus subulatus (Desv. ex Poir.) B.Øllg.
  • Phlegmariurus tafaensis (Nessel) A.R.Field & C.W.Chen
  • Phlegmariurus talamancanus (B.Øllg.) B.Øllg.
  • Phlegmariurus talamauanus (Alderw.) A.R.Field & Bostock
  • Phlegmariurus talpiphilus (B.Øllg.) B.Øllg.
  • Phlegmariurus tardieuae (Herter) A.R.Field & Testo
  • Phlegmariurus tauri (Herter) A.R.Field & Testo
  • Phlegmariurus taxifolius (Sm.) Á. & D.Löve
  • Phlegmariurus tenuicaulis (Underw. & F.E.Lloyd) B.Øllg.
  • Phlegmariurus tenuis (Humb. & Bonpl. ex Willd.) B.Øllg.
  • Phlegmariurus terrae-guilelmii (Herter) A.R.Field & Bostock
  • Phlegmariurus tetragonus (Hook. & Grev.) B.Øllg.
  • Phlegmariurus tetrastichoides (A.R.Field & Bostock) A.R.Field & Bostock
  • Phlegmariurus tetrastichus (Kunze) A.R.Field & Bostock
  • Phlegmariurus tico A.Rojas
  • Phlegmariurus toppingii (Herter) A.R.Field & Bostock
  • Phlegmariurus tournayanus Lawalrée
  • Phlegmariurus transilla (Sodiro ex Baker) B.Øllg.
  • Phlegmariurus treitubensis (Silveira) B.Øllg.
  • Phlegmariurus trifoliatus (Copel.) A.R.Field & Bostock
  • Phlegmariurus trigonus (C.Chr.) A.R.Field & Bostock
  • Phlegmariurus tryonorum B.Øllg.
  • Phlegmariurus tubulosus (Maxon) B.Øllg.
  • Phlegmariurus ulixis (Herter) B.Øllg.
  • Phlegmariurus unguiculatus (B.Øllg.) B.Øllg.
  • Phlegmariurus urbanii (Herter) B.Øllg.
  • Phlegmariurus vanuatuensis A.R.Field
  • Phlegmariurus varius (R.Br.) A.R.Field & Bostock
  • Phlegmariurus venezuelanicus (Herter) B.Øllg.
  • Phlegmariurus vernicosus (Hook. & Grev.) Á. & D.Löve
  • Phlegmariurus villonacensis (B.Øllg.) B.Øllg.
  • Phlegmariurus warneckei (Herter ex Nessel) A.R.Field & Testo
  • Phlegmariurus watsonianus (Maxon) B.Øllg.
  • Phlegmariurus weberbaueri (Nessel) B.Øllg.
  • Phlegmariurus weddellii (Herter) B.Øllg.
  • Phlegmariurus wilsonii (Underw. & F.E.Lloyd) B.Øllg.
  • Phlegmariurus xiphophyllus (Baker) A.R.Field & Bostock
  • Phlegmariurus yunfengii R.H.Jiang & X.C.Zhang
  • Phlegmariurus yunnanensis Ching

One hybrid is known:[1]

  • Phlegmariurus × koolauensis W.H.Wagner

References

  1. Hassler, Michael (19 January 2023), "Phlegmariurus", World Ferns. Synonymic Checklist and Distribution of Ferns and Lycophytes of the World, 14.7
  2. PPG I (2016), "A community-derived classification for extant lycophytes and ferns", Journal of Systematics and Evolution, 54 (6): 563–603, doi:10.1111/jse.12229, S2CID 39980610
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