Thick-billed fox sparrow

The megarhyncha subspecies group breeds in the mountains from southern Oregon to southern California east to the Sierra Nevada and shows little geographic variation. It interbreeds with the slate-colored subspecies group along a narrow contact zone from southern Oregon to western Nevada (Rising & Beadle 1996) but as noted above, gene flow is quite limited. Sibley (2000) indicates that this group has the most diagnostic call note, "a high, flat squeak [sic] teep like California towhee".

• megarhyncha Baird, 1858:

Breeds from the Onion Mountains and Robinson's Butte (southwestern Oregon) south through inland northern California to Kearsarge Pass (Inyo County), as well as the western flank of the Sierra Nevada in Mono County. Winters in central and southern California and adjacent Mexico.(Weckstein et al. 2002)

More larger-billed, duller, and grayer than the schistacea slate-colored fox sparrows; intermediate between the very large long-tailed stephensi and the more schistacea-like monoensis of this group; intergrades with the former.(Weckstein et al. 2002)

• stephensi Anthony, 1895:

Breeds in areas of the southern Sierra Nevada such as Fresno and Tulare Counties, Mount Pinos and the San Gabriel, San Bernardino, and San Jacinto Mountains, California, to the Sierra San Mártir, northern Baja California. In winter, migrates to lower elevations but generally does not migrate long distances. (Weckstein et al. 2002)

The largest, most massive-billed and longest-tailed fox sparrow; also very gray in coloration. (Swarth, 1920) Intergrades with megarhyncha. (Weckstein et al. 2002)

• monoensis Grinnell & Storer, 1917:

Breeds on the eastern flank of the Sierra Nevada in Mono County, California, and in the Walker River Range in adjacent Mineral County, Nevada. Winters from inland central California coastwards and south into Mexico. (Weckstein et al. 2002)

Somewhat intermediate between megarhyncha and schistacea slate-colored fox sparrows, with a paler, ashy gray back. (Weckstein et al. 2002)

• brevicauda Mailliard, 1918:

Breeds in the Mendocino Range of California, from the Yolla Bolly Mountains south to Snow Mountain and Sanhedrin Mountains. Migrates coastwards and south in winter. (Weckstein et al. 2002)

Similar to stephensi, but recognizable by more rusty tinge to brownish areas and the conspicuously shorter tail. (Weckstein et al. 2002)

• fulva Swarth, 1918:

Mountainous regions from central Oregon east of the Cascades Range (Steens Mountain, Sisters and Keeno Mountains) to Modoc and Lassen Counties, California. Winters in southwesternmost California and adjacent Mexico.

A browner version of monoensis; in winter range, distinguished from olivacea slate-colored fox sparrows by decidedly ruddy brown and larger bill. (Weckstein et al. 2002)

• mariposae Swarth, 1918:

Breeding in a limited area in California between the headwaters of the Little Shasta River (Siskiyou County) to Yosemite National Park and Kearsarge Pass. Winter range undocumented (Weckstein et al., 2002) due to lack of distinctiveness but probably same as megarhyncha and monoensis.

Probably nothing more than an intergrade between megarhyncha and monoensis; nowadays usually synonymized with the former. (Rising & Beadle 1996)

• Rising, Jim D. & Beadle, David (1996): A Guide to the Identification and Natural History of the Sparrows of the United States and Canada. Academic Press, San Diego. ISBN 0-12-588971-2
• Sibley, David Allen (2000): The Sibley Guide to Birds. Alfred A. Knopf, New York. ISBN 0-679-45122-6
• Swarth H. W. (1920): Revision of the avian genus Passerella with special reference to the distribution and migration of the races in California. University of California Publications in Zoology 21: 75–224.
• Weckstein, J. D.; Kroodsma, D. E. & Faucett, R. C. (2002): Fox Sparrow (Passerella iliaca). In: Poole, A. & Gill, F. (eds.): The Birds of North America 715. Academy of Natural Sciences, Philadelphia, PA & American Ornithologists' Union, Washington, D.C. Online version, retrieved 2006-11-27. doi:10.2173/bna.715 (requires subscription)
• Zink, R. M. (1994): The Geography of Mitochondrial DNA Variation, Population Structure, hybridization, and Species Limits in the Fox Sparrow (Passerella iliaca). Evolution 48(1): 96-111. doi:10.2307/2410006 (HTML abstract, first page image)
• Zink, Robert M. & Kessen, A. E. (1999): Species Limits in the Fox Sparrow. Birding 31: 508-517.
• Zink, Robert M. & Weckstein, Jason D. (2003): Recent evolutionary history of the Fox Sparrows (Genus: Passerella). Auk 120(2): 522–527. [Article in English with Spanish abstract] DOI: 10.1642/0004-8038(2003)120[0522:REHOTF]2.0.CO;2 HTML fulltext (without images)