Agraphydrus

Agraphydrus is a genus of water scavenger beetle in the family Hydrophilidae represented by 205 described species.[1] It is distributed across the Afrotropical, Australasian, and Indomalayan realms.

Agraphydrus
Dorsal view of Agraphydrussp.
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Family: Hydrophilidae
Subfamily: Acidocerinae
Genus: Agraphydrus
Régimbart, 1903
Diversity
205 species
Synonyms
  • Gymnhelochares d’Orchymont, 1932
  • Pseudohelochares Satô, 1960
  • Horelophopsis Hansen, 1997
  • Pseudopelthydrus Jia, 1998
  • Megagraphydrus Hansen, 1999

Taxonomy

The genus Agraphydrus was described for the first time by Régimbart in 1903.[2]

It currently contains 205 described species, partly thanks to a series revisionary work developed by Albretch Komarek.[3][4][5][6][7]

Description

Small beetles (1.4–4.8 mm), pale/yellowish to dark brown in coloration, eyes not emarginate, labrum exposed, maxillary palps moderately long. Elytra without sutural striae, not laterally explanate; the elytral punctation ranges from very fine and subtle to coarse.

A complete diagnosis was presented by Girón and Short,[1] but also see Komarek and collaborators.[3][4][5][6][7]

Habitat

According to Girón and Short,[1] based on annotations by Komarek and collaborators:[3][4][5][6][7]

Agraphydrus can be found in an extremely broad range of habitats, from rivers, streams and forest pools, to hygropetric environments around waterfalls or seepages over rocks; a few species have been collected in terrestrial habitats by sifting moss and leaves from near water bodies, or in the gravel along the bank of a river; in many cases specimens have been found associated with floating vegetation, mosses and algae.[1]

Species
  1. Agraphydrus abrasus Komarek and Freitag, 2020 [6]
  2. Agraphydrus activus Komarek and Hebauer, 2018 [7]
  3. Agraphydrus acutus Komarek, 2020 [5]
  4. Agraphydrus aethiopicus Komarek, 2020 [5]
  5. Agraphydrus agilis Komarek and Hebauer, 2018 [7]
  6. Agraphydrus albescens (Régimbart, 1903) [8]
  7. Agraphydrus ampullatus Komarek and Freitag, 2020 [6]
  8. Agraphydrus anacaenoides Komarek, 2019 [4]
  9. Agraphydrus anatinus Komarek, 2018 [3]
  10. Agraphydrus andamanicus Komarek, 2018 [3]
  11. Agraphydrus andringitra Komarek, 2020 [5]
  12. Agraphydrus angulatus Komarek, 2019 [4]
  13. Agraphydrus angustatus Komarek, 2020 [5]
  14. Agraphydrus angustipenis Komarek, 2018 [3]
  15. Agraphydrus anhuianus (Hebauer, 2000) [9]
  16. Agraphydrus annapurnensis Komarek, 2018 [3]
  17. Agraphydrus arduus Komarek and Hebauer, 2018 [7]
  18. Agraphydrus ater Komarek, 2018 [3]
  19. Agraphydrus atripalpis Komarek, 2020 [5]
  20. Agraphydrus attenuatus (Hansen, 1999) [10]
  21. Agraphydrus audax Komarek and Hebauer, 2018 [7]
  22. Agraphydrus avita (Hansen, 1997) [11]
  23. Agraphydrus bacchusi Komarek, 2019 [4]
  24. Agraphydrus balkeorum Komarek, 2019 [4]
  25. Agraphydrus batak Komarek and Freitag, 2020 [6]
  26. Agraphydrus bhutanensis Komarek, 2018 [3]
  27. Agraphydrus bicoloratus Komarek, 2020 [5]
  28. Agraphydrus bilardoi Komarek, 2020 [5]
  29. Agraphydrus biltoni Komarek, 2020 [5]
  30. Agraphydrus biprojectus Minoshima, Komarek, and Ôhara, 2015 [12]
  31. Agraphydrus borneensis Komarek, 2019 [4]
  32. Agraphydrus boukali Komarek, 2018 [3]
  33. Agraphydrus brevilobatus Komarek and Freitag, 2020 [6]
  34. Agraphydrus brevipenis Komarek, 2019 [4]
  35. Agraphydrus burmensis Komarek, 2019 [4]
  36. Agraphydrus calvus Komarek and Hebauer, 2018 [7]
  37. Agraphydrus camerunensis Komarek, 2020 [5]
  38. Agraphydrus cantonensis Komarek and Hebauer, 2018 [7]
  39. Agraphydrus carinatulus Komarek, 2019 [4]
  40. Agraphydrus cervus Komarek, 2019 [4]
  41. Agraphydrus ceylonensis Komarek, 2018 [3]
  42. Agraphydrus chinensis Komarek and Hebauer, 2018 [7]
  43. Agraphydrus cinnamum Komarek, 2018 [3]
  44. Agraphydrus clarus Komarek, 2019 [4]
  45. Agraphydrus comes Komarek and Hebauer, 2018 [7]
  46. Agraphydrus communis Komarek, 2018 [3]
  47. Agraphydrus confusus Komarek and Hebauer, 2018 [7]
  48. Agraphydrus congolensis Komarek, 2020 [5]
  49. Agraphydrus conicus Komarek and Hebauer, 2018 [7]
  50. Agraphydrus connexus Komarek and Hebauer, 2018 [7]
  51. Agraphydrus constrictus Komarek, 2018 [3]
  52. Agraphydrus contractus Komarek and Hebauer, 2018 [7]
  53. Agraphydrus coomani (d'Orchymont, 1927) [13]
  54. Agraphydrus coronarius Minoshima, Komarek, and Ôhara, 2015 [12]
  55. Agraphydrus crassipenis Komarek, 2018 [3]
  56. Agraphydrus dapengensis Yang & Jia, 2021 [14]
  57. Agraphydrus decipiens Minoshima, Komarek, and Ôhara, 2015 [12]
  58. Agraphydrus delineatus Komarek, 2019 [4]
  59. Agraphydrus elongatus Ribera, Hernando, and Cieslak, 2019 [15]
  60. Agraphydrus engkari Komarek, 2019 [4]
  61. Agraphydrus excisus Komarek, 2019 [4]
  62. Agraphydrus exedis (d'Orchymont, 1937) [16]
  63. Agraphydrus exiguus Komarek, 2019 [4]
  64. Agraphydrus falcatus Komarek, 2018 [3]
  65. Agraphydrus fasciatus Komarek and Hebauer, 2018 [7]
  66. Agraphydrus fikaceki Komarek and Hebauer, 2018 [7]
  67. Agraphydrus flavescens Komarek, 2020 [5]
  68. Agraphydrus flavipes Komarek, 2020 [5]
  69. Agraphydrus flavonotus Komarek, 2018 [3]
  70. Agraphydrus floresinus Komarek, 2019 [4]
  71. Agraphydrus fontis Komarek, 2020 [5]
  72. Agraphydrus forcipatus Komarek and Hebauer, 2018 [7]
  73. Agraphydrus fortis Komarek, 2018 [3]
  74. Agraphydrus fujianensis Komarek and Hebauer, 2018 [7]
  75. Agraphydrus geminus (d'Orchymont, 1932) [17]
  76. Agraphydrus gereckei Komarek, 2020 [5]
  77. Agraphydrus gilvus Komarek, 2018 [3]
  78. Agraphydrus glaber Komarek, 2018 [3]
  79. Agraphydrus globipenis Komarek and Hebauer, 2018 [7]
  80. Agraphydrus goldschmidti Komarek, 2020 [5]
  81. Agraphydrus gracilipalpis Komarek, 2018 [3]
  82. Agraphydrus hamatus Komarek, 2019 [4]
  83. Agraphydrus hanseni (Satô and Yoshitomi, 2004) [18]
  84. Agraphydrus heinrichi Komarek, 2018 [3]
  85. Agraphydrus helicopter Komarek, 2019 [4]
  86. Agraphydrus hendrichi Komarek, 2019 [4]
  87. Agraphydrus heterochromatus Komarek, 2019 [4]
  88. Agraphydrus hortensis Komarek, 2019 [4]
  89. Agraphydrus hygropetricus Komarek, 2018 [3]
  90. Agraphydrus igneus Komarek and Hebauer, 2018 [7]
  91. Agraphydrus imitans Komarek, 2019 [4]
  92. Agraphydrus indicus (d'Orchymont, 1932) [17]
  93. Agraphydrus inflatus Komarek, 2018 [3]
  94. Agraphydrus infuscatus Komarek, 2019 [4]
  95. Agraphydrus insidiator Minoshima, Komarek, and Ôhara, 2015 [12]
  96. Agraphydrus ishiharai (Matsui, 1994) [19]
  97. Agraphydrus jaechi (Hansen, 1999) [10]
  98. Agraphydrus jankodadai Komarek, 2019 [4]
  99. Agraphydrus jilanzhui Komarek and Hebauer, 2018 [7]
  100. Agraphydrus kallar Komarek, 2018 [3]
  101. Agraphydrus kathapa Komarek, 2019 [4]
  102. Agraphydrus kempi (d'Orchymont, 1922)
  103. Agraphydrus khasiensis Komarek, 2018 [3]
  104. Agraphydrus kodaguensis Komarek, 2018 [3]
  105. Agraphydrus komareki Yang & Jia, 2021 [14]
  106. Agraphydrus laocaiensis Komarek, 2019 [4]
  107. Agraphydrus latus Komarek, 2019 [4]
  108. Agraphydrus longipalpus (Jia, 1998) [20]
  109. Agraphydrus longipenis Komarek and Hebauer, 2018 [7]
  110. Agraphydrus lunaris Komarek, 2019 [4]
  111. Agraphydrus luteilateralis (Minoshima and Fujiwara, 2009) [21]
  112. Agraphydrus madagascarensis Komarek, 2020 [5]
  113. Agraphydrus maehongsonensis Komarek, 2019 [4]
  114. Agraphydrus malayanus (Hebauer, 2000) [9]
  115. Agraphydrus malkini Komarek, 2020 [5]
  116. Agraphydrus manfredjaechi Komarek, 2019 [4]
  117. Agraphydrus masatakai Minoshima, Komarek, and Ôhara, 2015 [12]
  118. Agraphydrus matoposensis Komarek, 2020 [5]
  119. Agraphydrus mazzoldii Komarek, 2019 [4]
  120. Agraphydrus meghalayanus Komarek, 2018 [3]
  121. Agraphydrus microphthalmus Komarek, 2019 [4]
  122. Agraphydrus minutissimus (Kuwert, 1890) [22]
  123. Agraphydrus mirabilis Komarek, 2019 [4]
  124. Agraphydrus montanus Minoshima, Komarek, and Ôhara, 2015 [12]
  125. Agraphydrus muluensis Komarek, 2019 [4]
  126. Agraphydrus musculus Komarek, 2019 [4]
  127. Agraphydrus namthaensis Komarek, 2019 [4]
  128. Agraphydrus nanus Komarek, 2018 [3]
  129. Agraphydrus narusei (Satô, 1960) [23]
  130. Agraphydrus nemorosus Komarek, 2019 [4]
  131. Agraphydrus nepalensis Komarek, 2018 [3]
  132. Agraphydrus niger Komarek, 2018 [3]
  133. Agraphydrus nigroflavus Komarek, 2019 [4]
  134. Agraphydrus obesus Komarek, 2019 [4]
  135. Agraphydrus obscuratus Komarek, 2018 [3]
  136. Agraphydrus obsoletus Komarek, 2018 [3]
  137. Agraphydrus occultus Komarek and Freitag, 2020 [6]
  138. Agraphydrus ogatai Minoshima, 2016 [24]
  139. Agraphydrus orbicularis Komarek, 2019 [4]
  140. Agraphydrus orientalis (d'Orchymont, 1932) [17]
  141. Agraphydrus palawanensis Komarek and Freitag, 2020 [6]
  142. Agraphydrus pallidus Komarek, 2019 [4]
  143. Agraphydrus papuanus Komarek, 2019 [4]
  144. Agraphydrus pauculus (Knisch, 1924) [25]
  145. Agraphydrus pauper Komarek, 2020 [5]
  146. Agraphydrus pelingeni Komarek and Freitag, 2020 [6]
  147. Agraphydrus penangensis Komarek, 2019 [4]
  148. Agraphydrus piceus Komarek, 2019 [4]
  149. Agraphydrus politus (Hansen, 1999) [10]
  150. Agraphydrus praecipuus (d'Orchymont, 1937) [26]
  151. Agraphydrus protentus Komarek, 2018 [3]
  152. Agraphydrus pseudoniger Yang & Jia, 2021 [14]
  153. Agraphydrus pullus Komarek, 2018 [3]
  154. Agraphydrus punctatellus Régimbart, 1903 [2]
  155. Agraphydrus punctulatus Komarek, 2018 [3]
  156. Agraphydrus puzhelongi (Jia, 2010) [27]
  157. Agraphydrus pygmaeus (Knisch, 1924) [25]
  158. Agraphydrus raucus Komarek, 2019 [4]
  159. Agraphydrus reductus Komarek and Hebauer, 2018 [7]
  160. Agraphydrus regularis (Hansen, 1999) [10]
  161. Agraphydrus reticulatus Komarek, 2019 [4]
  162. Agraphydrus reticuliceps Komarek and Hebauer, 2018 [7]
  163. Agraphydrus rhodesiensis Komarek, 2020 [5]
  164. Agraphydrus rhomboideus Komarek, 2019 [4]
  165. Agraphydrus rivalis Komarek, 2020 [5]
  166. Agraphydrus robustus Komarek and Hebauer, 2018 [7]
  167. Agraphydrus rostratus Komarek, 2018 [3]
  168. Agraphydrus rugosus Komarek, 2018 [3]
  169. Agraphydrus sabulosus Yang & Jia, 2021 [14]
  170. Agraphydrus sarawakensis Komarek, 2019 [4]
  171. Agraphydrus schoedli Komarek, 2019 [4]
  172. Agraphydrus schoenmanni Komarek and Hebauer, 2018 [7]
  173. Agraphydrus scintillans Komarek, 2019 [4]
  174. Agraphydrus scutifer Komarek, 2020 [5]
  175. Agraphydrus setifer Komarek and Hebauer, 2018 [7]
  176. Agraphydrus shaverdoae Komarek, 2019 [4]
  177. Agraphydrus siamensis (Hansen, 1999) [10]
  178. Agraphydrus sipekorum Komarek, 2018 [3]
  179. Agraphydrus skalei Komarek, 2019 [4]
  180. Agraphydrus spadix Komarek, 2019 [4]
  181. Agraphydrus spinosus Komarek, 2019 [4]
  182. Agraphydrus splendens Komarek and Hebauer, 2018 [7]
  183. Agraphydrus stagnalis (d'Orchymont, 1937)
  184. Agraphydrus stramineus Komarek, 2019 [4]
  185. Agraphydrus sucineus Komarek, 2019 [4]
  186. Agraphydrus sundaicus Komarek, 2019 [4]
  187. Agraphydrus tamdao Komarek, 2019 [4]
  188. Agraphydrus taprobanensis Komarek, 2018 [3]
  189. Agraphydrus tenuipalpis Komarek and Freitag, 2020 [6]
  190. Agraphydrus thaiensis Minoshima, Komarek, and Ôhara, 2015 [12]
  191. Agraphydrus tristis Komarek, 2019 [4]
  192. Agraphydrus tulipa Komarek, 2019 [4]
  193. Agraphydrus tumidus Komarek, 2020 [5]
  194. Agraphydrus tumulosus Komarek, 2018 [3]
  195. Agraphydrus umbrosus Komarek and Hebauer, 2018 [7]
  196. Agraphydrus uncinatus Komarek and Hebauer, 2018 [7]
  197. Agraphydrus usambaraensis Komarek, 2020 [5]
  198. Agraphydrus uvaensis (Hebauer, 2000) [9]
  199. Agraphydrus vadoni Komarek, 2020 [5]
  200. Agraphydrus variabilis Komarek and Hebauer, 2018 [7]
  201. Agraphydrus vietnamensis Komarek, 2019 [4]
  202. Agraphydrus villiersi (Balfour-Browne, 1958) [28]
  203. Agraphydrus wangmiaoi Komarek and Hebauer, 2018 [7]
  204. Agraphydrus yunnanensis Komarek and Hebauer, 2018 [7]
  205. Agraphydrus zetteli Komarek and Freitag, 2020 [6]

References
  1. Girón, Jennifer C.; Short, Andrew Edward Z. (2021-06-18). "The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species". ZooKeys (1045): 1–236. doi:10.3897/zookeys.1045.63810. ISSN 1313-2970. PMC 8233300. PMID 34228772.
  2. Régimbart, MA (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France. 72: 1–51.
  3. Komarek, A. (2018). "Taxonomic revision of AgraphydrusRégimbart, 1903 II. The Indian Subcontinent" (PDF). Koleopterologische Rundschau. 88: 103–204. Archived from the original (PDF) on 2021-06-24. Retrieved 2022-01-07.
  4. Komarek, A. (2019). "Taxonomic revision of AgraphydrusRégimbart, 1903 III. Southeast Asia (except Philippines) and Australian Region (Coleoptera: Hydrophilidae: Acidocerinae)" (PDF). Koleopterologische Rundschau. 89: 151–316. Archived from the original (PDF) on 2021-06-24. Retrieved 2022-01-07.
  5. Komarek, A. (2020). "Taxonomic revision of AgraphydrusRégimbart, IV. Africa, Western Asia, and redescription of the genus (Coleoptera: Hydrophilidae: Acidocerinae)" (PDF). Koleopterologische Rundschau. 90: 127–200. Archived from the original (PDF) on 2021-02-14. Retrieved 2022-01-07.
  6. Komarek, A.; Freitag, H. (2020). "Taxonomic revision of AgraphydrusRégimbart, 1903 V. Philippine species and their first DNA barcodes" (PDF). Koleopterologische Rundschau. 90: 201–242. Archived from the original (PDF) on 2021-02-14. Retrieved 2022-01-07.
  7. Komarek, A.; Hebauer, F. (2018-07-30). "Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)". Zootaxa. 4452 (1): 1–101. doi:10.11646/zootaxa.4452.1.1. ISSN 1175-5334. PMID 30314249. S2CID 52975164.
  8. Régimbart, M. (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophi-lidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France. 72: 1–51.
  9. Hebauer, F. (2000). "The genus Megagraphydrus Hansen, 1999, with description of new species (Coleoptera: Hydrophilidae)". Acta Coleopterologica. 16: 14–22.
  10. Hansen, Michael (1999-01-01). "Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family". Insect Systematics & Evolution. 30 (2): 121–172. doi:10.1163/187631200X00228. ISSN 1876-312X.
  11. Hansen, M. (1997). "A new subfamily for a remarkable new genus and species of Hydrophilidae from New Guinea (Coleoptera: Hydrophilidae)". Annales Zoologici. 47: 107–110.
  12. Minoshima, Y. N.; Komarek, A.; Ôhara, M. (2015). "A revision of Megagraphydrus Hansen (Coleoptera, Hydrophilidae): synonymization with Agraphydrus Régimbart and description of seven new species". Zootaxa. 3930: 1–63. PMID 25781810.
  13. d’Orchymont, A. (1927). "Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums". Journal of the Federated Malay States Museums. 13 (4): 246–252.
  14. Yang, Zhen-ming; Jia, Fenglong; Jiang, Lu; Guo, Qiang (2021-05-18). "Four new species of Agraphydrus Régimbart, 1903 with additional faunistic record from China (Coleoptera, Hydrophilidae, Acidocerinae)". Deutsche Entomologische Zeitschrift. 68 (1): 189–205. doi:10.3897/dez.68.66200. ISSN 1860-1324. S2CID 236330170.
  15. Ribera, I.; Hernando, C.; Cieslak, A. (2019). "Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae" (PDF). Acta Entomologica Musei Nationalis Pragae. 59: 253–272. doi:10.2478/aemnp-2019-0021. hdl:10230/44839. S2CID 198381025.
  16. d’Orchymont, A. (1937). "Descriptions of three new Hydrophilidae from India". Records of the Indian Museum. 39: 29–31.
  17. d’Orchymont, A. (1932). "Zur Kenntnis der Kolbenwasserkäfer (Palpicornia) von Sumatra, Java und Bali". Archiv für Hydrobiologie. Supplement Band IX: 623–714. [pl. XIV–XVII.]
  18. Satô, M.; Yoshitomi, H. (2004). "Discovery of a second representative of the genus Horelophopsis (Coleoptera, Hydrophilidae) from the Ryukyu archipelago, Japan". Elytra. 32: 41–49.
  19. Matsui, E. (1994). "Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society. 20: 215–220.
  20. Jia, F. (1998). "A new genus Pseudopelthydrus gen. n. from Hainan Island, China (Coleoptera: Hydrophilidae: Hydrophilinae)". Chinese Journal of Entomology. 18: 225–230.
  21. Minoshima, Y. N.; Fujiwara, J. (2009). "Discovery of water scavenger beetle genus Megagraphydrus Hansen (Coleoptera, Hydrophilidae) from Japan, with description of a new species" (PDF). Elytra. 37: 53–64.
  22. Kuwert, A. (1890). "Bestimmungs-Tabellen der europäischen Coleopteren. XX. Heft. Hydrophilidae. II. Abteilung: Sphaeridiini und Helophorini". Verhandlungen des naturforschenden Vereins in Brünn. 28: 159–328.
  23. Satô, M. (1960). "One new genus and two new species of the subtribe Helocharae from Japan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society. 6: 76–80.
  24. Minoshima, Yûsuke N. (2016). "Taxonomic review of Agraphydrus from Japan (Coleoptera: Hydrophilidae: Acidocerinae)". Entomological Science. 19 (4): 351–366. doi:10.1111/ens.12213. ISSN 1479-8298. S2CID 89394202.
  25. Knisch, A. (1924). "Neue neotropische Palpicornier. (Col. Hydrophilidae. - Op. 16.)". Wiener Entomologische Zeitung. 41: 114–140.
  26. d’Orchymont, A. (1937). "Contribution à l'étude des Palpicornia IX". Bulletin et Annales de la Société Entomologique de Belgique. 77: 213–255.
  27. Jia, Fenglong (2010-06-08). "Megagraphydrus puzhelongi, sp. n., a new water scavenger beetle from China (Coleoptera: Hydrophilidae: Hydrophilinae)". Zootaxa. 2498 (1): 65. doi:10.11646/zootaxa.2498.1.6. ISSN 1175-5334.
  28. Balfour-Browne, J. (1958). "La Reserve naturelle integrale du Mont Nimba, Guinea. VI. Coléoptères Hydrophiloides". Mémoires de l'Institut français d'Afrique noire. 53: 169–190.
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