Aguja Formation
The Aguja Formation is a geological formation in North America, exposed in Texas, United States and Chihuahua and Coahuila in Mexico, whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.[1] Fossil palms have also been unearthed here.[2]
Aguja Formation | |
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Stratigraphic range: Lower to Middle Campanian ~ | |
![]() Outcrops of the Aguja Formation, Big Bend National Park, Texas | |
Type | Geological formation |
Unit of | Tornillo Group |
Sub-units | Upper Shale, Terlingua Creek Sandstone, Rattlesnake Mt. Sandstone, Lower Shale & Basal Sandstone Members |
Underlies | Javelina Formation |
Overlies | Pen Formation |
Lithology | |
Primary | Sandstone, conglomerate, claystone |
Other | Mudstone, shale, limestone |
Location | |
Coordinates | 29.3°N 103.5°W |
Approximate paleocoordinates | 35.8°N 77.0°W |
Region | ![]() ![]() ![]() |
Country | ![]() ![]() |
![]() ![]() Aguja Formation (the United States) ![]() ![]() Aguja Formation (Texas) |
![](../I/North_america_75mya.png.webp)
Age
The ages of the Aguja Formation and its primary fossil-bearing unit, the Upper Shale, are not well understood. Due to the presence of the ammonite Baculites mclearni, which only occurs from 80.67 - 80.21 Ma, in the underlying Rattlesnake Mountain Sandstone and the Terlingua Creek Sandstone, it is likely that the Upper Shale was younger than 80.2 Ma.[3] A radiometric date of 76.9 Ma was recovered in the Upper Shale, making it likely the formation wasn't younger than 76.9 Ma.[3] The contact with the overlying Javelina Formation has been estimated at about 70 Ma ago[4] but also as recently as 68.5 million years ago.[5] This is unlikely, however, due to the presence of Bravoceratops, more primitive than an unnamed chasmosaurine from the De-na-zin Member of the Kirtland Formation, in the lowermost section of the formation.[6] The age of the Basal Sandstone is constrained by the presence of Scaphites hippocrepis III in the overlying Pen Formation which has been dated as old as 81.53 Ma.[3][7]
Paleofauna
![](../I/Hadrosaur_bone_in_the_Aguja_Formation_by_Nick_Longrich.jpg.webp)
Reptiles
2 fragmentary caudal vertebrae of indeterminate reptiles are known from the Rattlesnake Mountain sandstone member.[8]
Pseudosuchians
Crurotarsans of the Aguja Formation | ||||||
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Genus | Species | Location | Member | Material | Notes | Images |
Crocodilia | Indeterminate | Ten Bits Ranch, west Texas.[8] | Rattlesnake Mountain sandstone member.[8] | Teeth.[8] | A crocodilian. | |
Deinosuchus[9] | D. riograndensis | Texas,[9] Chihuahua, and Coahuila[10] | Osteoderm and mandible fragment | A giant alligatoroid. | ![]() | |
cf. D. sp. | West Texas[11] | Lower Shale[11] | Cervical osteoderm (TMM 44068-2).[11] | An alligatoroid. | ![]() | |
Goniopholididae | New genus & species | West Texas[11] | Lower Shale[11] | Partial skull & skeleton along with isolated osteoderms & teeth.[11] | May pertain to Denazinosuchus or a related taxon. | |
Phobosuchus[9] | P. riograndensis[9] | Texas,[9] Chihuahua, and Coahuila | Reclassified as a Deinosuchus species | |||
Ornithischians
Ornithischians of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Agujaceratops | A. mariscalensis | Texas[12] | Low Upper Shale[13] | Formerly considered a species of Chasmosaurus[12] | ![]() | |
A. mavericus | West Texas[14] | A chasmosaurine. | ||||
Angulomastacator | A. daviesi | Texas[15] | Upper Shale[15] | A left maxilla.[15] | A lambeosaurine. | ![]() |
Ankylosauridae | Indeterminate | Texas[16] | Osteoderms, vertebrae & limb elements.[16] | Remains of an ankylosaurid, possibly represents Euoplocephalus sp.[16] | ||
Aquilarhinus[17] | A. palimentus | Texas[18] | Lower Shale[18] | Formerly referred to Kritosaurus and Gryposaurus.[19] | ![]() | |
Ceratopsidae genus & species indeterminate | Indeterminate | West Texas[11] | Lower Shale[11] | Ilia, sacral vertebra & sacral ribs.[11] | May represent Agujaceratops, but undiagnostic. | |
Chasmosaurus[20] | C. mariscalensis[21] | Texas[21] | [Twelve] disarticulated skull (sic), postcrania, juvenile."[22] | Considered by paleontologists Lucas, Sullivan, and Hunt to be distinct enough from the Chasmosaurus type species, C. belli to warrant being split off to a new genus, Agujaceratops.[12] | ||
Edmontonia | E. sp[23] | Texas[16] | Osteoderm (TVP 45866-2) & skull (AMNH 3076).[16] | A nodosaurid. | ![]() | |
cf. Euoplocephalus[21] | E. sp[21] | Texas[16] | Upper Shale[16] | Osteoderms, sacrum & vertebra (TL-05-14).[16] | An ankylosaurid. | ![]() |
Kritosaurus[20] | Indeterminate[20] | Chihuahua[24] | ![]() | |||
Malefica | M. deckerti | Texas[25] | Upper Shale[25] | A partial left maxilla.[25] | Formerly referred to Kritosaurus.[25] | |
Nodosauridae genus & species indeterminate | Indeterminate | West Texas[11] | Lower Shale[11] | Isolated & associated osteoderms.[11] | Very similar to osteoderms of Invictarx. | |
Panoplosaurus | P. mirus | Texas[16] | Osteoderm (TMM 45605-4).[16] | A nodosaurid. | ![]() | |
Texacephale | T. langstoni | Texas[13] | Low Upper Shale[13] | Two frontoparietal domes.[13] | A pachycephalosaur. | ![]() |
Yehuecauhceratops | Y. mudei[26] | Coahuila[27] | A centrosaurine. | ![]() | ||
Theropods
Indeterminate ornithomimid remains are known from the Upper Aguja Formation.[28] Indeterminate tyrannosaurid fossils are known from the Upper Aguja Formation of Texas and Mexico.[29]
Theropods of the Aguja Formation | ||||||
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Genus | Species | Location | Member | Material | Notes | Images |
Avialae incertae sedis | West Texas[30] | Lower Shale[30] | 4 teeth (TMM 45947-349, 350, 351).[30] | A bird. | ||
Chirostenotes | C. sp. | Texas[31] | Femur & manual ungual.[31] | A caenagnathid | ![]() | |
Dromaeosauridae genus & species indeterminate | Indeterminate | West Texas[11] | Lower Shale[11] | Pedal phalanges (TMM 45909-2, TMM 44066-4).[11] | A dromaeosaur. | |
cf. Dromaeosaurus[32] | Indeterminate[32] | Texas[21] | ![]() | |||
Leptorhynchos | L. gaddisi | Texas[31] | Dentaries, caudal vertebra & limb elements.[31] | A caenagnathid | ![]() | |
Ornithomimidae | New genus & species | West Texas[11] | Lower Shale[11] | Vertebrae, ischium & limb elements.[11] | An unnamed species referred to informally as the 'Aguja ornithomimid'. | |
cf. Paronychodon | West Texas[30] | Lower Shale[30] | 2 incomplete teeth (TMM 45947-362).[30] | A maniraptoran. | ![]() | |
Richardoestesia | R. cf. gilmorei | Texas[33] | Low Upper Shale[33] | Fragment of a small tooth.[33] | A coelurosaur. | ![]() |
R. isosceles | Big Bend National Park, Texas[33] | Low Upper Shale[33] | Teeth.[33] | A coelurosaur. | ||
Saurornitholestes | S. cf. langstoni | Texas[30][33] | Low Upper Shale & Lower Shale[30][33] | Teeth.[30][33] | A dromaeosaur. | ![]() |
Theropoda incertae sedis | Morphotype A | West Texas[30] | Lower Shale[30] | 8 teeth.[30] | Unserrated, recurved teeth. | |
Morphotype B | West Texas[30] | Lower Shale[30] | 5 tooth crowns.[30] | Bi-serrated teeth. | ||
Morphotype C | West Texas[30] | Lower Shale[30] | 7 teeth.[30] | Finely serrated, distal plication only. | ||
Morphotype D | West Texas[30] | Lower Shale[30] | 5 teeth.[30] | Short, coarsely serrated distal plication. | ||
Morphotype E | West Texas[30] | Lower Shale[30] | A tooth.[30] | Strongly recurved, serrated distal plication. | ||
Morphotype F | West Texas[30] | Lower Shale[30] | 2 tooth fragments.[30] | Medium size, finely serrated plications. | ||
cf. Troodon[34] | Indeterminate[34] | Texas[21] | ![]() | |||
Tyrannosauridae | Indeterminate | West Texas[30][35] | Upper Shale[35] & Lower Shale[11][30] | Isolated teeth & a handful of non-dental specimens.[30][35] | A relatively small & gracile tyrannosaurid.[35] | |
Lepidosaurs
Lepidosaurs of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Anguidae | Gen. et. sp. indet. | Texas.[36] | Isolated osteoderms & partial right frontal.[36] | An anguid. | ||
Apsgnathus | A. triptodon | Brewster County, Texas.[36] | Jaw elements.[36] | A scincomorph. | ||
Catactegenys | C. solaster | Brewster County, Texas.[36] | Jaw elements & teeth.[36] | A night lizard. | ||
Dryadissector | D. shilleri | West Texas.[30] | Lower Shale.[30] | Numerous isolated teeth.[30] | A varanoid. | |
Mosasauridae | Indeterminate | Ten Bits Ranch.[8] | Rattlesnake Mountain sandstone member.[8] | 1 partial vetebra.[8] | A mosasaur. | |
Odaxosaurus | O. piger | Brewster County, Texas.[36] | Jaw elements.[36] | An anguid. | ||
cf. Parasaniwa | cf. P. wyomingensis | Texas.[36] | Jaw fragments.[36] | A platynotan. | ||
Platynota | Gen. et. sp. indet. | Texas.[36] | Dorsal vertebra (TMM 43057-332).[36] | A platynotan. | ||
?Scincidae | Gen. et. sp. indet. | Brewster County, Texas.[36] | Jaw elements.[36] | A possible skink. | ||
cf. Scincomorpha | Gen. et. sp. indet. | Brewster County, Texas.[36] | Jaw elements.[36] | A scincomorph. | ||
Serpentes | Gen. et. sp. indet. | Texas.[36] | Partial left dentary & right maxilla.[36] | A snake. | ||
cf. Xenosauridae | Texas.[36] | Osteoderms & maxillae.[36] | A knob-scaled lizard. | |||
Turtles
Testudines of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Adocus | A. sp. | Big Bend National Park, Texas.[37] | Upper Shale[37] | Shell fragments.[37] | An adocid. | ![]() |
Baenidae | Big Bend National Park, Texas.[37] | Upper Shale[37] | Shell fragments.[37] | 2 morphotypes present (A & B). | ||
Basilemys | B. sp. | Big Bend National Park, Texas.[37] | Upper Shale & Lower Shale[11][37] | Shell fragments, partial plastron & leg scutes.[11][37] | A nanhsiungchelyid. | ![]() |
Bothremydidae genus & species indeterminate | Indeterminate | West Texas[11] | Lower Shale[11] | Isolated peripheral bones (TMM 44068-1, 42452-8); costal bones (TMM 44064-6).[11] | A bothremydid, compatible with Chupacabrachelys. | |
Chupacabrachelys | C. complexus | Big Bend, Texas.[38] | Base of the Upper Shale[38] | A complete skull, and a nearly complete skeleton.[38] | A bothremydid. | ![]() |
cf. Denazinemys | cf. D. sp. | West Texas[11] | Lower Shale[11] | Shell elements.[11] | A baenid. | |
cf. Helopanoplia | Big Bend National Park, Texas.[37] | Upper Shale[37] | Shell fragments.[37] | A softshell turtle. | ||
cf. Hoplochelys | Big Bend National Park, Texas.[37] | Upper Shale[37] | Shell fragments.[37] | A kinosternoid. | ||
Terlinguachelys | T. fischbecki | Big Bend National Park, Texas[39] | Rattlesnake Mountain Sandstone[39] | A large, incomplete specimen.[39] | A protostegid. | |
Testudines indeterminate[8] | Morphotype 1[8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | A shell fragment. | ||
Morphotype 2[8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | A shell fragment. | |||
Morphotype 3[8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | A shell fragment. | |||
Morphotype 4[8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | A shell fragment. | |||
Trionychidae | Gen. et. sp. indet. | Big Bend National Park, Texas.[37] | Upper Shale & Lower Shale[11][37] | Costal bone (TMM 44068-4) & shell fragments.[37] | A softshell turtle. | |
Bony Fish
Approximately 75 whole and broken fragments of coprolites are known from the Rattlesnake Mountain sandstone member, presumably from bony fish.[8]
Bony fish of the Aguja Formation | ||||||
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Genus | Species | Location | Member | Material | Notes | Images |
Albula | A. sp. | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Over 390 complete and fragmentary teeth.[40] | A bonefish. | ![]() |
?Enchodus | ?E. sp. | Ten Bits ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 2 abraded teeth.[40] | An aulopiform. | ![]() |
Eotexachara | E. malateres | West Texas[41] | Dentaries.[41] | A characiform. | ||
Lepidotes | ?L. sp. | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Approximately 109 complete and fragmentary teeth.[40] | A lepidotid. | |
Melvius | M. sp. | West Texas[11] | Lower Shale[11] | Vertebrae.[11] | An amiid. | |
Osteichthyes | Indeterminate species A | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 11 complete teeth. | ||
Indeterminate species B | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 8 complete and fragmentary teeth. | |||
Indeterminate species C | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 1 complete tooth. | |||
Indeterminate species D | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 1 complete and 1 partial tooth. | |||
Indeterminate | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | Approximately 475 complete and fragmentary teleost centra.[40] | |||
Paralbula | P. casei | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Approximately 900 complete and fragmentary teeth and tooth caps.[40] | ||
Primuluchara | P. laramidensis | West Texas[41] | Dentaries.[41] | A characiform. | ||
Stephanodus | ?S. sp. | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 21 whole and fragmentary specimens.[40] | A pycnodont. | |
Cartilaginous fish
Cartilaginous fish of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Material | Notes | Images |
Brachyrhyzodus | B. wichitaensis | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 11 complete teeth.[40] | ||
Cantioscyllium | C. aff. meyeri | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 1 anterior tooth and 5 lateral teeth.[8] | A nurse shark. | |
Chiloscyllium | C. aff. greeni | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8] | Approximately 90 abraded and fragmentary teeth.[40] | A bamboo shark. | |
Chondrichthyes[8] | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 23 placoid scales and 69 dermal scales. | 4 morphotypes of placoid scales (A to D) present. | ||
Columbusia | C. sp. | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 20 complete & fragmentary teeth.[40] | A wobbegong. | |
Cretalamna | C. appendiculata | Ten Bits Ranch, west Texas.[8] | Rattlesnake Mountain sandstone member.[8] | 3 fragmentary teeth.[8] | Reassigned to C. cf. C. sarcoportheta.[40] | |
C. cf. C. sarcoportheta | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 3 incomplete teeth & fragments of additional teeth.[40] | Originally reported as C. appendiculata. | ![]() | |
Cretorectolobus[8] | C. olsoni | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | "20 complete and fragmentary teeth". | A carpet shark. | |
Hybodontidae genus & species indeterminate | Indeterminate | West Texas[11] | Lower Shale[11] | Fragment of a dorsal fin spine (TMM 42536-10).[11] | A hybodont. | |
Hybodus[8] | H. sp. | Ten Bits Ranch. | Rattlesnake Mountain sandstone member.[8] | 2 specimens, one complete and one partial tooth. | A hybodont. | ![]() |
Igdabatis | I. indicus? | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | A single incomplete tooth.[8][40] | A myliobatid. | |
Ischyrhiza | I. cf. avonicola | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 5 complete and fragmentary rostral teeth.[40] | A sawskate. | |
I. mira | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 11 fragmentary rostral teeth and 230 whole and fragmentary oral teeth.[40] | A sawskate. | ![]() | |
Lonchidion | L. selachos | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 4 complete and 5 fragmentary teeth.[8][40] | A hybodont. | |
Meristodon | M. sp. | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 2 teeth.[40] | A hybodont. | |
Myliobatiformes | Incertae sedis | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 3 complete specimens.[40] | ||
Protoplatyrhina | P. renae | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8] | 70 complete and fragmentary teeth.[40] | A hypsobatid. | |
Ptychotrygon | P. agujaensis | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Over 690 complete and fragmentary teeth.[40] | A sawskate. | |
P. triangularis | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Over 170 complete and fragmentary teeth.[40] | A sawskate. | ||
P. aff. cuspidata | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | 7 whole and fragmentary teeth.[40] | A sawskate. | ||
P. sp. | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | One complete tooth (TMM 46018-71).[40] | A sawskate. | ![]() | |
Rhinobatos | R. casieri | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | "14 complete and fragmentary specimens". | A guitarfish. | |
R. sp. | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 18 complete & fragmentary specimens.[40] | A guitarfish. | ||
Rhombodus | R. levis | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Over 22 complete and fragmentary teeth.[40] | A rajiforme. | |
Scapanorhynchus | S. texanus | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | Over 800 complete and fragmentary teeth.[40] | A mitsukurinid. | ![]() |
Sclerorhynchidae | Morphotype 1 | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[8] | One fragmentary rostral spine (TMM 46018-59).[40] | A sawskate. | |
Morphotype 2 | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[8] | One fragmentary rostral spine (TMM 46018-60).[40] | A sawskate. | ||
Morphotype 3 | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[8] | 3 fragmentary rostral teeth.[40] | A sawskate. | ||
Morphotype 4 | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[8] | 27 fragmentary rostral spines.[40] | A sawskate. | ||
Serratolamna | S. cf. S. caraibaea | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | About 34 teeth.[40] | A mackerel shark. | |
Squalicorax | S. kaupi | Ten Bits Ranch, west Texas.[8][40] | Rattlesnake Mountain sandstone member.[8][40] | "26 complete and fragmentary anterior and lateral teeth". | An anacoracid. | ![]() |
S. aff. S. lindstromi | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 4 specimens.[40] | An anacoracid. | ||
S. pristodontus | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 2 specimens. | An anacoracid. | ![]() | |
S. aff. S. yangaensis | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 4 complete & several fragmentary teeth.[40] | An anacoracid. | ||
Squatina | S. hassei | Ten Bits Ranch, west Texas.[8] | Rattlesnake Mountain sandstone member.[8] | A single complete specimen. | An angelshark. | |
S. sp. | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | A complete tooth & 2 fragmentary teeth.[40] | An angelshark. | ||
Texatrygon | T. cf. T. copei | Ten Bits Ranch, west Texas.[40] | Rattlesnake Mountain sandstone member.[40] | 4 complete & fragmentary teeth.[40] | A sawskate formerly reported as T. hooveri. | |
T. hooveri | Ten Bits Ranch, west Texas. | Rattlesnake Mountain sandstone member.[8] | 4 complete and fragmentary specimens.[8] | Reassigned to T. cf. T. copei. |
Ammonites
Ammonites of the Aguja Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Member | Abundance | Notes | Images |
Baculites | B. mclearni |
|
||||
Hoplitoplacenticeras | H. plasticum | Rattlesnake Mountain Sandstone | ||||
Pachydiscus | P. paulsoni | Rattlesnake Mountain Sandstone | ![]() | |||
Color key
|
Notes Uncertain or tentative taxa are in small text; |
References
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- Fowler, Denver Warwick (2017-11-22). "Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America". PLOS ONE. 12 (11): e0188426. Bibcode:2017PLoSO..1288426F. doi:10.1371/journal.pone.0188426. ISSN 1932-6203. PMC 5699823. PMID 29166406.
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- "Aguja Formation (Upper Shale Member), Big Bend Region, Texas," in Sullivan and Lucas (2006). Page 16.
- "The first mandible fragment of Deinosuchus (Eusuchia: Alligatoroidea) discovered in Coahuila, Mexico". ResearchGate. Retrieved 2016-03-27.
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- West, Bryanna (2020). Campanian-Maastrictian Ankylosaurs of West Texas (PDF) (Thesis). Texas Tech University.
- Listed as ?Gryposaurus sp. in "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
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- "Dinosaur distribution (Texas and Chihuahua)." Weishampel, et al. (2004). Page 582 and 588.
- "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
- "Table 23.1," in Weishampel, et al. (2004). Page 496.
- Listed as Edmontonia cf. rugosidens in "Dinosaur distribution (Late Cretaceous; North America; Texas).", Weishampel et al., 2004, p.582
- "Dinosaur distribution (Late Cretaceous; North America; Chihuahua)." Weishampel, et al. (2004). Page 588.
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- Rivera-Sylva, H.E.; Frey, E.; Stinnesbeck, W.; Guzman-Gutirrez, J.R.; Gonzalez-Gonzalez (2017). "Mexican ceratopsids: Considerations on their diversity and evolution". Journal of South American Earth Sciences. doi:10.1016/j.jsames.2017.01.008.
- Sullivan, R.M., and Lucas, S.G. 2006. "The Kirtlandian land-vertebrate "age" – faunal composition, temporal position and biostratigraphic correlation in the nonmarine Upper Cretaceous of western North America." New Mexico Museum of Natural History and Science, Bulletin 35:7-29.
- Mortimer, M (2004). "Tyrannosauroidea". The Theropod Database. Archived from the original on 2013-09-29. Retrieved 2007-08-21.
- Wick, Steven L.; Lehman, Thomas M.; Brink, Alyson A. (2015). "A theropod tooth assemblage from the lower Aguja Formation (early Campanian) of West Texas, and the roles of small theropod and varanoid lizard mesopredators in a tropical predator guild". Palaeogeography, Palaeoclimatology, Palaeoecology. 418: 229–244. Bibcode:2015PPP...418..229W. doi:10.1016/j.palaeo.2014.11.018. ISSN 0031-0182.
- Longrich, Nicholas R.; Barnes, Ken; Clark, Scott; Millar, Larry (April 2013). "Caenagnathidae from the Upper Campanian Aguja Formation of West Texas, and a Revision of the Caenagnathinae". Bulletin of the Peabody Museum of Natural History. 54 (1): 23–49. doi:10.3374/014.054.0102. ISSN 0079-032X. S2CID 128444961.
- Listed as cf. Dromaeosaurus sp. in "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
- Sankey, Julia T. (2001). "Late Campanian Southern Dinosaurs, Aguja Formation, Big Bend, Texas". Journal of Paleontology. 75 (1): 208–215. doi:10.1666/0022-3360(2001)075<0208:LCSDAF>2.0.CO;2. ISSN 0022-3360. JSTOR 1306931.
- Listed as cf. Troodon sp. in "Dinosaur distribution (Late Cretaceous; North America; Texas)." Weishampel, et al. (2004). Page 582.
- Lehman, Thomas M.; Wick, Steven L. (September 2012). "Tyrannosauroid dinosaurs from the Aguja Formation (Upper Cretaceous) of Big Bend National Park, Texas". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 471–485. doi:10.1017/S1755691013000261. ISSN 1755-6910. S2CID 129232391.
- Nydam, Randall L.; Rowe, Timothy B.; Cifelli, Richard L. (2013). "Lizards and Snakes of the Terlingua Local Fauna (late Campanian), Aguja Formation, Texas, with Comments on the Distribution of Paracontemporaneous Squamates Throughout the Western Interior of North America". Journal of Vertebrate Paleontology. 33 (5): 1081–1099. Bibcode:2013JVPal..33.1081N. doi:10.1080/02724634.2013.760467. ISSN 0272-4634. JSTOR 42568627. S2CID 86519841.
- Sankey, Julia (January 2006). "Turtles of the upper Aguja Formation (late Campanian), Big Bend National Park, Texas". New Mexico Museum of Natural History and Science Bulletin. 35: 235–243.
- Lehman, Thomas M.; Wick, Steven L. (2010). "Chupacabrachelys complexus, N. Gen. N. Sp. (testudines: Botheremydidae), from the Aguja Formation (campanian) of West Texas". Journal of Vertebrate Paleontology. 30 (6): 1709–1725. Bibcode:2010JVPal..30.1709L. doi:10.1080/02724634.2010.520782. ISSN 0272-4634. JSTOR 25790797. S2CID 129303574.
- Lehman, Thomas M.; Tomlinson, Susan L. (November 2004). "Terlinguachelys fischbecki, a new genus and species of sea turtle (Chelonioidea: Protostegidae) from the Upper Cretaceous of Texas". Journal of Paleontology. 78 (6): 1163–1178. doi:10.1666/0022-3360(2004)078<1163:TFANGA>2.0.CO;2. ISSN 0022-3360. S2CID 54755373.
- Schubert, Joseph A.; Wick, Steven L.; Lehman, Thomas M. (January 2017). "An Upper Cretaceous (middle Campanian) marine chondrichthyan and osteichthyan fauna from the Rattlesnake Mountain sandstone member of the Aguja Formation in West Texas". Cretaceous Research. 69: 6–33. Bibcode:2017CrRes..69....6S. doi:10.1016/j.cretres.2016.08.008. S2CID 133098369.
- Wick, Steven L. (2021-12-01). "New early Campanian characiform fishes (Otophysi: Characiformes) from West Texas support a South American origin for known Late Cretaceous characiforms from North America". Cretaceous Research. 128: 104993. Bibcode:2021CrRes.12804993W. doi:10.1016/j.cretres.2021.104993. ISSN 0195-6671.
Bibliography
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