Ardeadactylus

Ardeadactylus (from Ardea – meaning "heron", and also a name of a genus of herons – and dactylus, meaning "finger") is an extinct genus of ctenochasmatoid pterosaur known from the Late Jurassic Solnhofen limestone of Bavaria, southern Germany. It contains a single species, Ardeadactylus longicollum, which was originally thought to be a species of Pterodactylus, as P. longicollum.[1]

Ardeadactylus
Temporal range: Late Jurassic,
SMNS 56603, the neotype
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Clade: Euctenochasmatia
Clade: Ctenochasmatoidea
Genus: Ardeadactylus
Bennett, 2013
Species:
A. longicollum
Binomial name
Ardeadactylus longicollum
Synonyms
List
  • Pterodactylus longipes
    Münster, 1836
  • Pterodactylus longicollum
    Meyer, 1854
  • Diopecephalus longicollum
    (Meyer, 1854)
  • Pterodactylus (Ornithocephalus) vulturinus
    Wagner, 1857
  • Pterodactylus vulturinus
    (Wagner, 1857)
  • Pterodactylus suevicus
    Fraas, 1878 non Quenstedt, 1855
  • Cycnorhamphus fraasi
    Seeley, 1901
  • Pterodactylus fraasi
    (Seeley, 1901)

History

Only two specimens of Ardeadactylus are known to exist currently: SMNS 56603 (earlier SMNS 5802) found in 1874, a specimen from Nusplingen initially thought to belong to the species Pterodactylus suevicus (currently Cycnorhamphus),[2] and the neotype of the species, JME-SOS 2428, a specimen held at Jura Museum in Eichstätt. Other known specimens, including the holotype designated by Christian Erich Hermann von Meyer when he named the type species Pterodactylus longicollum in 1854,[3] were lost during World War II.[1] The original holotype, consisting of the skull, neck and anterior torso, had been found near Eichstätt in 1853 and was that year acquired for the Herzoglich Leuchtenbergische Naturalien-Kabinett by Professor Ludwig Frischmann.[3] Its destruction motivated Peter Wellnhofer to assign a neotype in 1970.[4]

Description

Ardeadactylus is suspected to be a heron–like, long–necked long–legged piscivore.[1] It was similar to Pterodactylus antiquus in its general body form, but seems to have been larger. Bennett (2013) estimated the wingspan of the neotype specimen to be 1.45 meters (4.8 ft); the referred specimen from the Jura Museum is more or less 10% larger.[1] A. longicollum had fewer, fifteen per jaw, and relatively larger teeth than P. antiquus, possibly indicating that it preyed on larger fish than Pterodactylus.[1]

Classification

Illustration of the lost holotype from 1860

In 2017, a phylogenetic analysis by Steven Vidovic and David Martill found Ardeadactylus within the family Aurorazhdarchidae, sister taxon to Aurorazhdarcho. Their cladogram is shown below:[5]

Ctenochasmatoidea
Aurorazhdarchia

Aerodactylus

Aurorazhdarchidae

Huanhepterus

Ardeadactylus

Aurorazhdarcho

Gallodactylidae

Cycnorhamphus

Moganopterinae

Feilongus

Moganopterus

Ctenochasmatidae

In 2018 however, Nicholas Longrich and colleagues had recovered a different set of relationships for ctenochasmatoids. In their analysis, Ardeadactylus was recovered within the Ctenochasmatidae instead. Their cladogram is shown below.[6]

Ctenochasmatidae
Ctenochasmatinae

Ctenochasma

Pterodaustrini

Pterodaustro

Beipiaopterus

Gegepterus

Kepodactylus

Moganopterinae

Moganopterus

Feilongus

Ardeadactylus

Elanodactylus

Gnathosaurinae

Huanhepterus

Plataleorhynchus

Gnathosaurus

See also

References

  1. Bennett, S. C. (2013). "New information on body size and cranial display structures of Pterodactylus antiquus, with a revision of the genus". Paläontologische Zeitschrift. 87 (2): 269–289. doi:10.1007/s12542-012-0159-8. S2CID 83722829.
  2. Fraas, O., 1878, "Über Pterodactylus suevicus, Qu., von Nusplingen", Palaeontographica, 25: 163–174
  3. Meyer, C.E.H. von, 1854, "Mittheilungen an Professor Bronn: Anthracotherium Dalmatinum vom Monte Promina u.a. A.; Chelydra Decheni aus Braunkohle des Siebengebirges; Wirbelthier-Reste aus dem Basalttuff-Konglomerat zu Glimbach an der Rabenau; angebliches Vorkommen von Agnotherium antiquum und Hyaena spelaea; fossile Reste im lithographischen Schiefer von Nusplingen bei Spaichingen; Eryon Schuberti; Litogaster; Pemphix; Pterodactylus longicollum n. sp. in Solenhofener Schiefer; Acrosaurus Frischmanni von da; Reptilien und Cancer-Arten im Kressenberger Nummuliten-Gestein", Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde 1854: 47–58
  4. Wellnhofer, P., 1970, "Die Pterodactyloidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands", Bayerische Akademie der Wissenschaften, Mathematisch-Wissenschaftlichen Klasse, Abhandlungen 141: 1–133
  5. Vidovic, Steven U.; Martill, David M. (2017). "The taxonomy and phylogeny of Diopecephalus kochi (Wagner, 1837) and "Germanodactylus rhamphastinus" (Wagner, 1851)" (PDF). Geological Society, London, Special Publications. 455 (1): 125–147. Bibcode:2018GSLSP.455..125V. doi:10.1144/SP455.12. S2CID 219204038.
  6. Longrich, N.R.; Martill, D.M.; Andres, B. (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3): e2001663. doi:10.1371/journal.pbio.2001663. PMC 5849296. PMID 29534059.
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