Arizonasaurus

Arizonasaurus was a ctenosauriscid archosaur from the Middle Triassic (243 million years ago).[1] Arizonasaurus is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, Arizonasaurus babbitti, was named by Samuel Paul Welles in 1947.

Arizonasaurus
Temporal range: Middle Triassic (Anisian), Possible Carnian record
Model, Museum am Löwentor, Stuttgart
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pseudosuchia
Clade: Paracrocodylomorpha
Superfamily: †Poposauroidea
Family: †Ctenosauriscidae
Genus: †Arizonasaurus
Welles, 1947
Species
  • †A. babbitti Welles, 1947 (type)

Discovery and naming

The type species, Arizonasaurus babbitti, was named by Samuel Paul Welles in 1947 on the basis of a few teeth and a maxilla, labelled as specimen UCMP 36232.[2] A fairly complete skeleton was found in 2002 by Sterling Nesbitt.[3]

Description
Restoration

Arizonasaurus had a sail made of tall neural spines. This sail was similar to those of other basal archosaurs, such as other ctenosauriscids like Ctenosauriscus, Lotosaurus, Bromsgroveia, and Hypselorhachis.[4]

Arizonasaurus is described from two braincase specimens. Some ancestral features of these braincases are plesiomorphic for crurotarsans.[1]

Below is a list of characteristics found by Nesbitt in 2005 that distinguish Arizonasaurus:[4]

  • a deep fossa hidden from view on the posteroventral edge of the upward-pointing process of the maxilla;
  • and a tongue-groove attachment between the pubis and the ilium.

Classification
Fossil

Arizonasaurus was closely related to Ctenosauriscus; and, together with a few other genera, they make up Ctenosauriscidae. The ctenosauriscids were closely related to the poposaurids, as shown by a few shared derived characteristics.[3] The pelvic girdle in Arizonasaurus unites this taxon with Ctenosauriscus, Lotosaurus, Bromsgroveia, and Hypselorhachus.[4] Together, newly identified pseudosuchian features act as evidence that poposaurids, such as Poposaurus, Sillosuchus, and Chatterjeea, and ctenosauriscids form a monophyletic group that is derived rauisuchians.[3]

Below is a phylogenetic cladogram simplified from Butler et al. in 2011 showing the cladistics of Archosauriformes, focusing mostly on Pseudosuchia:[5]

Poposauroidea

Qianosuchus

Ctenosauriscidae

Arizonasaurus

Xilousuchus

Hypselorhachis

Ctenosauriscus

Bromsgroveia

Waldhaus Taxon

Poposaurus gracilis H

Poposaurus gracilis Y

Lotosaurus

Sillosuchus

Shuvosaurus

Effigia

Biogeography

Arizonasaurus is from the middle Triassic Moenkopi Formation of northern Arizona. The presence of a poposauroid in the early Middle Triassic suggests that the divergence of birds and crocodiles occurred earlier than previously thought. Ctenosauriscids from the Middle Triassic allow the distribution of Triassic faunas to be more widespread, now in Europe, Asia, North America and Africa. The fauna of the Moenkopi Formation represents a stage transitional fauna between the faunas of older and younger age.[3]

References
  1. Gower, D.J. & Nesbitt, S.J. (2006). "The braincase of Arizonasaurus babbitti-further evidence for the non-monophyly of 'rauisuchian' archosaurs". Journal of Vertebrate Paleontology. 26 (1): 79–87. doi:10.1671/0272-4634(2006)26[79:TBOABE]2.0.CO;2. S2CID 55338555.
  2. Welles, S.P. (1947). "Vertebrates from the Upper Moenkopi Formation of northern Arizona". University of California Publications in Geological Sciences. 27 (7): 241–294.
  3. Nesbitt, S.J. (2003). "Arizonasaurus and its implications for archosaur divergence". Proceedings of the Royal Society B. 270: S234–S237. doi:10.1098/rsbl.2003.0066. PMC 1809943. PMID 14667392.
  4. Nesbitt, S.J. (2005). "Osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti". Historical Biology. 8 (1): 19–47. doi:10.1080/08912960500476499. S2CID 84326151.
  5. Butler, R.J.; Brusatte, S.J.; Reich, M.; Nesbitt, S.J.; Schoch, R.R.; Hornung, J.J. (2011). Andrew A. Farke (ed.). "The Sail-Backed Reptile Ctenosauriscus from the Latest Early Triassic of Germany and the Timing and Biogeography of the Early Archosaur Radiation". PLOS ONE. 6 (10): e25693. Bibcode:2011PLoSO...625693B. doi:10.1371/journal.pone.0025693. PMC 3194824. PMID 22022431.
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