Benthophilinae

Benthophilinae
	The pale monkey goby (Neogobius fluviatilis) and the dark bighead goby (Ponticola kessleri) are representatives of Benthophilinae
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Gobiiformes
Family:	Gobiidae
Subfamily:	Benthophilinae; Beling & Iljin, 1927
Type genus
	Benthophilus; Eichwald, 1831
Genera
	See text

The Benthophilinae are a subfamily of gobies endemic to the Ponto-Caspian region (including the Marmara, Black, Azov, Caspian, and Aral Seas).[1] The subfamily includes about 50 species. The representatives of the subfamily have fused pelvic fins and elongated dorsal and anal fins.[2] They are distinguished from the closely related subfamily Gobiinae by the absence of a swimbladder in adults and location of the uppermost rays of the pectoral fins within the fin membrane.[3]

The Catalog of Fishes still considers these fishes as belonging to the subfamily Gobiinae.[4]

Systematics of the subfamily

Tribe Benthophilini
- Anatirostrum
- Benthophiloides
- Benthophilus (type genus)
- Caspiosoma
Tribe Neogobiini
- Neogobius (type genus)
Tribe Ponticolini
- Babka
- Mesogobius
- Ponticola (type genus)
- Proterorhinus[5]

References

Simonović P.D., Nikolić V.P., Skóra K.E. (1996) Vertebral number in Ponto-Caspian gobies: phylogenetic relevance. J. Fish Biol., 49: 1027–1029.
Miller P.J. (1986) Gobiidae. In: Whitehead P.J.P., Bauchot M.-L., Hureau J.-C., Nielsen J., Tortonese E. (eds.) Fishes of the North-eastern Atlantic and the Mediterranean, Vol. 3. UNESCO, Paris.
Pinchuk V.I. (1991) K voprosu o grupirovkakh vidov v predelakh roda Neogobius (Perciformes). Voprosy Ikhtiologii, 31: 380–393.
W. Eschmeyer (2015) Catalog of Fishes Archived May 3, 2015, at the Wayback Machine California Academy of Sciences (1 Mar 2015)
Neilson M.E., Stepien C.A. (2009) Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei). Molecular Phylogenetics and Evolution, 52(1): 84-102.

External links

Neogobiin research at the University of Toledo, Ohio
Systematics of the Neogobiin Gobies Research project at U Toledo

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[1] Simonović P.D., Nikolić V.P., Skóra K.E. (1996) Vertebral number in Ponto-Caspian gobies: phylogenetic relevance. J. Fish Biol., 49: 1027–1029.

[2] Miller P.J. (1986) Gobiidae. In: Whitehead P.J.P., Bauchot M.-L., Hureau J.-C., Nielsen J., Tortonese E. (eds.) Fishes of the North-eastern Atlantic and the Mediterranean, Vol. 3. UNESCO, Paris.

[3] Pinchuk V.I. (1991) K voprosu o grupirovkakh vidov v predelakh roda Neogobius (Perciformes). Voprosy Ikhtiologii, 31: 380–393.

[4] W. Eschmeyer (2015) Catalog of Fishes Archived May 3, 2015, at the Wayback Machine California Academy of Sciences (1 Mar 2015)

[5] Neilson M.E., Stepien C.A. (2009) Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei). Molecular Phylogenetics and Evolution, 52(1): 84-102.