Obrimini

The Obrimini differ from their sister tribe the Hoplocloniini by the structure of the secondary ovipositor at the abdomen of the females, which surrounds the actual ovipositor. It is formed in the Obrimini dorsally from the eleventh abdominal tergum called supraanal plate or epiproct, while in the Hoplocloniini it originated from the tenth tergum.[2][4][5]

The distribution area of the Obrimini extends from Borneo to the east and includes the Philippines, Sulawesi, most of the Moluccas and New Guinea. Farthest east lies with Viti Levu, the main island of the Fiji group, the distribution area of Pterobrimus depressus.[2][5]

Brunner von Wattenwyl built in 1893 for the genera already described Obrimus, Hoploclonia, Tisamenus, Pylaemenes, Dares and Datames (today a synonym to Pylaemenes) the tribe Obrimini (abbreviated there as Obrimi.). He placed these together with the genus Heteropteryx and the South American tribe Cladomorphini (abbreviated there as Cladomorphi.) in the family Cladomorphidae (now synonymous with Cladomorphinae).[6] In the following years, basesd on this work, mostly genera of today's Obriminae and Dataminae were listed in this tribe, for example by Josef Redtenbacher in 1906.[4] Lawrence Bruner raised the Obrimini to the rank of a family in 1915. James Abram Garfield Rehn and his son John William Holman Rehn divided the Obriminae, which they only referred to as a subfamily, into the tribes Obrimini and Datamini in 1939.[7] Both tribes were transferred in 1953 by Klaus Günther to the subfamily Heteropteryginae.[8] In 2004 Oliver Zompro raised this subfamily to the rank of family and the tribes contained in the rank of subfamilies or in the rank of a separate family (Anisacanthidae). At the same time he built three tribes in the subfamily Obriminae. In addition to the Obrimini, these were the Miroceramiini and the Eubulidini.[9] The latter was in 2016 by Frank H. Hennemann et al synonymized with the Obrimini. At the same time, the new tribe Tisamenini was established within the Obriminae.[5] In a study based on genetic analysis, published in 2021, Sarah Bank et al synonymized the tribe Miroceramiini and the tribe Tisamenini with the Obrimini and placed next to this the Hoplocloniini as the only other tribe in the subfamily Obriminae. In addition, a species previously referred to as Trachyaretaon sp. 'Negros' was identified as a representative of an as yet undescribed genus.[2]

Genera

Image	Genus	Species
	Aretaon Rehn, J.A.G. & Rehn, J.W.H., 1939	Aretaon asperrimus (Redtenbacher, 1906) Aretaon muscosus (Redtenbacher, 1906)
	Brasidas Rehn, J.A.G. & Rehn, J.W.H., 1939	Brasidas acanthoderus Rehn, J.A.G. & Rehn, J.W.H., 1939 Brasidas foveolatus (Redtenbacher, 1906) Brasidas montivagus Rehn, J.A.G. & Rehn, J.W.H., 1939 Brasidas quadratipes (Bolívar, 1890) Brasidas samarensis Rehn, J.A.G. & Rehn, J.W.H., 1939 Brasidas viscayanus Rehn, J.A.G. & Rehn, J.W.H., 1939
	Eubulides Stål, 1877	Eubulides alutaceus Stål, 1877 Eubulides igorrote Rehn, J.A.G. & Rehn, J.W.H., 1939 Eubulides taylori Rehn, J.A.G. & Rehn, J.W.H., 1939
	Euobrimus Rehn, J.A.G. & Rehn, J.W.H., 1939	Euobrimus atherura Rehn, J.A.G. & Rehn, J.W.H., 1939 Euobrimus bakeri Rehn, J.A.G. & Rehn, J.W.H., 1939 Euobrimus cavernosus Stål, 1877 Euobrimus cleggi Rehn, J.A.G. & Rehn, J.W.H., 1939 Euobrimus dohrni Rehn, J.A.G. & Rehn, J.W.H., 1939 Euobrimus hoplites Rehn, J.A.G. & Rehn, J.W.H., 1939 Euobrimus lacerta (Redtenbacher, 1906) Euobrimus stephenreyesi Lit & Eusebio, 2006
	Heterocopus Redtenbacher, 1906	Heterocopus leprosus Redtenbacher, 1906
	Mearnsiana Rehn, J.A.G. & Rehn, J.W.H., 1939	Mearnsiana bullosa Rehn, J.A.G. & Rehn, J.W.H., 1939
	Miroceramia Günther, 1934	Miroceramia westwoodii (Bates, 1865)
	Obrimus Stål, 1875	Obrimus bicolanus Rehn, J.A.G. & Rehn, J.W.H., 1939 Obrimus bufo (Westwood, 1848) Obrimus mesoplatus (Westwood, 1848) Obrimus uichancoi Rehn, J.A.G. & Rehn, J.W.H., 1939
	Pterobrimus Redtenbacher, 1906	Pterobrimus depressus Redtenbacher, 1906
	Stenobrimus Redtenbacher, 1906	Stenobrimus bolivari Redtenbacher, 1906 Stenobrimus lumad Lit & Eusebio, 2010 Stenobrimus pilipinus Eusebio, Lit & Lucañas, 2023 Stenobrimus tagalog Rehn, J.A.G. & Rehn, J.W.H., 1939
	Sungaya Zompro, 1996	Sungaya inexpectata Zompro, 1996
	Theramenes Stål, 1875	Theramenes dromedarius Stål, 1877 Theramenes exiguus Hennemann & Conle, 2003 Theramenes letiranti Hennemann, 2023[10] Theramenes mandirigma Zompro & Eusebio, 2001 Theramenes olivaceus (Westwood, 1859)
	Tisamenus Stål, 1875	Tisamenus alviolanus Lit & Eusebio, 2010 Tisamenus armadillo Redtenbacher, 1906 Tisamenus asper Ignacio Bolívar, 1890 Tisamenus atropos (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus cervicornis Ignacio Bolívar, 1890 Tisamenus clotho (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus deplanatus (Westwood, 1848) Tisamenus draconina (Westwood, 1848) Tisamenus fratercula (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus hystrix (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus hebardi (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus kalahani Lit & Eusebio, 2005 Tisamenus lachesis (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus polillo (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus ranarius (Westwood, 1859) Tisamenus serratorius Stål, 1875 Tisamenus spadix (Rehn, J.A.G. & Rehn, J.W.H., 1939) Tisamenus summaleonilae Lit & Eusebio, 2005 Tisamenus tagalog (Rehn, J.A.G. & Rehn, J.W.H., 1939)
	Trachyaretaon Rehn, J.A.G. & Rehn, J.W.H., 1939	Trachyaretaon brueckneri Hennemann & Conle, 2006 Trachyaretaon carmelae Lit & Eusebio, 2005 Trachyaretaon echinatus (Stål, 1877) Trachyaretaon gatla Zompro, 2004

• Data related to Obrimini at Wikispecies