Paraná Formation

Paraná Formation
Paraná Formation
Stratigraphic range: Late Miocene
Type	Geological formation
Location
Region	Entre Ríos Province
Country	Argentina
Type section
Named for	Paraná Basin

The Paraná Formation is a geologic formation in Argentina. It preserves fossils dating back to the late Miocene epoch.

Fossil content

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Cetaceans

Cetaceans reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
cf. Aulophyseter	cf. A. sp.	Cantera del Puerto Viejo near Paraná, Entre Ríos.[1]	Incomplete tooth (MACN Pv 8926).[1]	Now deemed an indeterminate physeteroid.[2]
Balaenidae gen. et. sp. indet.	Indeterminate		Right tympanic bulla (MACN Pv 13221).[1]	A balaenid.
Balaenoptera	B. sp.		Right tympanic bulla (MACN Pv 13220).[1]	A rorqual.
Balaenopteridae indet.		South Bank of Ensenada Creek.[3]	Cubitus & radius (CICYTTP-PV-1-197) bearing bite marks matching Cosmopolitodus hastalis.[3]	A rorqual.
Physeteroidea gen. et. sp. indet.	Indeterminate	Cantera del Puerto Viejo near Paraná, Entre Ríos.[2]	An almost complete tooth (MAS Pv 1361) & a fragmentary tooth (MACN Pv 8926).[2]	Previously identified as cf. Aulophyseter sp.[2]
Platanistoidea indet.			Left tympanic bulla (MACN Pv 15997).[1]	A relative of river dolphins.
Pontistes	P. rectifrons		[4]	A pontoporiid.
Prionodelphis	P. rovereti		A single tooth.[5]	A dubious delphinoid, other specimens once referred to this taxon are now thought to represent monachine seals.[5]

Pinnipeds

Pinnipeds reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Properiptychus	P. argentinus		Jaw & limb elements.[6]	A monachine seal.

Rodents

Rodents reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
cf. Cardiatherium	cf. C. sp. indet.	La Juanita locality.[7]	Left isolated cheek tooth (CICYTTP-PV-M-1-147) & right upper cheek tooth (CICYTTP-PV-M-1-148).[7]	A hydrochoerine.
Caviidae gen. & sp. indet.	Indeterminate	La Juanita locality.[7]	A left upper M1-2 (CICYTTP-PV-M-2-267).[7]	A cavy.
Plesiacarechimys	P. nov. sp.?	La Juanita locality.[7]	Right lower molar (CICYTTP-PV-M-1-145).[7]	An octodontoid.

Sirenians

Sirenians reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Dioplotherium	D. sp.	Cliffs along Paraná River, Villa Urquiza.[8]	Upper left molar (CORD-PZ 4301).[8]	A dugongid.
Metaxytherium	M. sp.	Cerro La Matanza locality, Victoria City.[8]	Posterior end of both maxillae (MASP 373).[8]	A dugongid.

Xenarthrans

Xenarthrans reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Scirrotherium	S. sp. indet.	La Juanita locality.[7]	A fixed osteoderm (CICYTTP-PV-M-1-144).[7]	A pampatheriid.

Birds

Birds reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Pelecanus	P. paranensis	Cerro La Matanza locality, Victoria City.[9]	A nearly complete pelvis (CICYTTP-PV-A-3-277).[9]	A pelican.
cf. Phoenicopterus	cf. P. sp. indet.	La Juanita locality.[7]	Fragment of shaft with distal end of left tibiotarsus (CICYTTP-PV-A-1-12).[7]	A flamingo.

Bony fish

Bony fish reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Siluroidae	Undetermined		Abundant fragmentary specimens.[10]	Evidently represents extant catfish genera.[10]
Tetraodontidae indet.	Indeterminate	La Juanita & Toma Vieja localities.[11]	Several premaxillary beaks.[11]	Remains of pufferfish.

Rays

Rays reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Dynatobatis	D. paranensis		Dermal tubercles.[10]
Myliobatis	M. americanus		Detached teeth.[10]
Raja	R. agassizi		Dermal tubercles.[10]	Moved to the genus Rioraja.
Rioraja	R. agassizii		Dermal tubercles.[10]	A skate, originally reported as Raja agassizi.

Sharks

Sharks reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Carcharias	C. (Prionodon) obliquidens		Teeth.[10]	Junior synonym of Squalus obliquidens.[12]
Carcharodon	C. megalodon		Teeth.[10]	Reassigned to the genus Otodus.
Carcharodon	C. sp.	Arroyo Ensenada valley.[13]	2 teeth (UAP 1301 & 1303).[13]	Related to the great white shark.
Cestracion	C. paranensis		2 teeth.[10]	Cestracion is now a junior synyonym of Heterodontus.
Cosmopolitodus	C. hastalis		Several teeth.[10]	A mackerel shark, originally reported as Oxyrhina hastalis.
Galeocerdo	G. aduncus		Tooth.[10]	A requiem shark.
Hemipristis	H. serra		A few well-preserved upper teeth.[10]	A large weasel shark.
Heterodontus	H. paranensis		2 teeth.[10]	A bullhead shark, originally reported as Cestracion paranensis.
Megascyliorhinus	M. trelewensis	La Juanita locality.[14]	4 anterior teeth.[14]	A ground shark, specimens originally labelled "Squatina".[14]
Odontaspis	O. elegans		Teeth.[10]
Otodus	O. megalodon		Teeth.[10]	A megatooth shark, originally reported as Carcharodon megalodon.
Oxyrhina	O. hastalis		Several teeth.[10]	Reassigned to the genus Cosmopolitodus.
Squalus	S. obliquidens		Teeth.[10]	Originally reported as Carcharias (Prionodon) obliquidens.

Bivalves

Bivalves reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Aequipecten	A. paranensis		Valves.[15]	A scallop.
Amusium	A. darwinianum		[15]	A scallop.
Erodona	E. doellojuradoi	La Juanita & La Paz.[16]	260 specimens.[16]	An erodonid.
Flabellipecten	F. oblongus		Valves.[15]	A scallop.
Polymesoda	P. muravchiki	La Juanita.[16]	13 specimens.[16]	A cyrenid.

Bryozoans

Bryozoans reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Acanthodesia	A. cf. sulcata	Punta Gorda, near Diamante, Entre Ríos.[17]	Fragments of colonies (MLP 13471 to 13476).[17]	A cheilostomatid.
Microporella	M. sp.	Punta Gorda, near Diamante, Entre Ríos.[17]	Fragments of colonies (MLP 13477 to 13483).[17]	A microporellid.
Schizoporella	S. sp.	Punta Gorda, near Diamante, Entre Ríos.[17]	Fragments of colonies (MLP 13474, MLP 13484 to 13488).[17]	A schizoporellid.
Schizoporellidae indet.	Indeterminate	Punta Gorda, near Diamante, Entre Ríos.[17]	Fragments of colonies (MLP 13489).[17]	A schizoporellid.
Schizosmittina	S. sp.	Punta Gorda, near Diamante, Entre Ríos.[17]	Fragments of colonies (MLP 13486, MLP 13490 to 13492).[17]	A bitectiporid.
Tubulipora	T. sp.	Punta Gorda, near Diamante, Entre Ríos.[17]	Two colonies (MLP 13493 & MLP 13494).[17]	A tubuliporid.

Echinoderms

Echinoderms reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Abertella	A. sp.		[18]	A sand dollar.
Amplaster	A. alatus		[18]	A sand dollar.
Amplaster	A. ellipticus		[18]	A sand dollar.
Monophoraster	M. duboisi		[18]	A sand dollar.
Neognathostomata	Indeterminate		An immature specimen.[18]	A sea urchin.
Ophiocoma	O. sp.		[18]	A brittle star.
Ophiothrix	O. sp.		[18]	A brittle star.

Plants

Plants reported from the Paraná Formation
Genus	Species	Presence	Material	Notes	Images
Astroniumxylon	A. parabalansae	Toma Vieja fossil locality.[19]	Fossil wood.[19]	An anacardiaceaen.
Laurophyllum	L. sp.	Villa Urquiza.[20]	Leaves.[20]	A lauraceaen.
Myrciophyllum	M. paranaesianum	Villa Urquiza.[20]	Leaves.[20]	A myrtaceaen.
Ocotea	O. sp.	Villa Urquiza.[21]	Leaves.[21]	A lauraceaen.
Piptadenioxylon	P. paraexcelsa	Toma Vieja fossil locality.[19]	Fossil wood.[19]	A mimosoid.
Schinus	S. aff. terebinthifolius	Villa Urquiza.[20]	Leaves.[20]	An anacardiaceaen.
Solanumxylon	S. paranensis	Toma Vieja fossil locality.[19][22]	Fossil wood.[19]	A nightshade.

Correlations

Laventan

Laventan correlations in South America
Formation	Honda	Honda	Aisol	Cura-Mallín	Pisco	Ipururo	Pebas	Capadare	Inés	Paraná	Map
Basin	VSM	Honda	San Rafael	Caldera	Pisco	Ucayali	Amazon	Falcón	Venezuela	Paraná	Paraná Formation (South America)
Country	Colombia	Bolivia	Argentina	Chile	Peru			Venezuela		Argentina
Boreostemma
Hapalops
Miocochilius
Theosodon
Xenastrapotherium
Mylodontidae
Sparassodonta
Primates
Rodents
Birds
Terror birds
Reptiles
megalodon
Flora
Insects
Environments	Fluvial			Fluvio-deltaic		Fluvio-lacustrine		Fluvio-deltaic		Fluvial	Laventan volcanoclastics Laventan fauna Laventan flora
Volcanic	Yes										Laventan volcanoclastics Laventan fauna Laventan flora

Huayquerian

Huayquerian correlations in South America
Formation	Cerro Azul	Ituzaingó	Camacho	Raigón	Andalhuala	Las Flores	Maimará	Palo	Pebas	Muyu	Rosa	Saldungaray	Salicas	Urumaco	Map
Basin	Colorado	Paraná			Hualfín	Tontal	Andes	Salta	Amazon	Huasi	Altiplano	BA	Velasco	Falcón	Paraná Formation (South America)
Country	Argentina		Uruguay		Argentina				Brazil Peru	Bolivia		Argentina		Venezuela
Cardiatherium
Lagostomus
Macroeuphractus
Proeuphractus
Pronothrotherium
Pseudotypotherium
Thylacosmilus
Xotodon
Macraucheniidae
Primates
Rodents
Reptiles
Birds
Terror birds
Flora
Environments	Aeolian-fluvial	Fluvio-deltaic		Fluvial					Fluvio-lacustrine	Fluvial			Fluvio-lacustrine	Fluvio-deltaic	Huayquerian volcanoclastics Huayquerian fauna Huayquerian flora
Volcanic					Yes

References

Agnolin, Federico; Lucero, Sergio (December 2004). "Registros de cetáceos del Mioceno Tardío de Entre Ríos, Argentina". Revista de biología marina y oceanografía. 39 (2). doi:10.4067/S0718-19572004000200007. ISSN 0718-1957.
Pérez, Leandro M.; Cione, Alberto L.; Cozzuol, Mario; Varela, Augusto N. (June 2011). "A Sperm Whale (Cetacea: Physeteroidea) from the Paraná Formation (Late Miocene) of Entre Rís, Argentina. Environment and Taphonomy". Ameghiniana. 48 (4): 648–654. doi:10.5710/AMGH.v48i3(425). ISSN 0002-7014. S2CID 129118367.
Noriega, Jorge Ignacio; Cione, Alberto Luis; Aceñolaza, Florencio Gilberto (2007-09-05). "Shark tooth marks on Miocene balaenopterid cetacean bones from Argentina". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 245 (2): 185–192. doi:10.1127/0077-7749/2007/0245-0185. ISSN 0077-7749.
Cozzuol, Mario A. (April 1996). "The record of aquatic mammals in southern South America". Münchner Geowiss. Abh. 30: 321–342.
Muizon, Christian de; Hendey, Q. B. (1980). "Late Tertiary seals of the south Atlantic Ocean". Annals of the South African Museum. Annale van die Suid-Afrikaanse Museum. 82: 91–128. ISSN 0303-2515.
de Muizon, Christian; Bond, Mariano (1982). "Le Phocidae (Mammalia) de la Formation Parana (Entre Rios, Argentine)". Bull. Mus. Natn. Hist. Nat., Paris (in French): 165–207.
Candela, Adriana M.; Bonini, Ricardo A.; Noriega, Jorge I. (November 2012). "First continental vertebrates from the marine Paraná Formation (Late Miocene, Mesopotamia, Argentina): Chronology, biogeography, and paleoenvironments". Geobios. 45 (6): 515–526. Bibcode:2012Geobi..45..515C. doi:10.1016/j.geobios.2012.05.003. S2CID 128431992.
Vélez-Juarbe, Jorge; Noriega, Jorge I.; Ferrero, Brenda S. (2012). "FOSSIL DUGONGIDAE (MAMMALIA, SIRENIA) FROM THE PARANÁ FORMATION (LATE MIOCENE) OF ENTRE RÍOS PROVINCE, ARGENTINA". Ameghiniana. 49 (4): 585–593. doi:10.5710/AMGH.27.6.2012.568. hdl:11336/72358. ISSN 0002-7014. S2CID 56435219.
Noriega, Jorge I.; Cenizo, Marcos; Brandoni, Diego; Pérez, Leandro M.; Tineo, David E.; Diederle, Juan M.; Bona, Paula (2023-05-09). "A new pelican (Aves: Pelecanidae) from the Upper Miocene of Argentina: new clues about the origin of the New World lineages". Journal of Vertebrate Paleontology. doi:10.1080/02724634.2023.2202702. ISSN 0272-4634. S2CID 258605013.
Woodward, Arthur Smith (1900). "I.—On some fish-remains from the Parana formation, Argentine Republic". The Annals and Magazine of Natural History; Zoology, Botany, and Geology. 6 (31): 1–7. doi:10.1080/00222930008678334. ISSN 0374-5481.
"Tetraodontiformes (Osteichthyes, Actinopterygii) from the Miocene of Argentina: with the southernmost record of fossil Tetraodontidae". Comptes Rendus Palevol. 20 (27): 585–596. 2021.
"Squalus obliquidens | Shark-References". shark-references.com. Retrieved 2023-05-21.
Cione, Alberto Luis; Cabrera, Daniel Alfredo; Barla, María Julia (2012-03-01). "Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic". Geobios. 45 (2): 167–172. Bibcode:2012Geobi..45..167C. doi:10.1016/j.geobios.2011.06.002. ISSN 0016-6995.
Cione, Alberto Luis; Mennucci, Jorge; P&eacute, Leandro; rez; Marí Barla, a Julia (2008-01-01). "Megascyliorhinus trelewensis (Neoselachii) in the Middle-Upper Miocene of Parana, Central Eastern Argentina". Instituto Superior de Correlacion Geologica, Serie Miscelanea. 17 (1): 41–49.
Martín Pérez, Leandro; Griffin, Miguel; Iturerría, Santiago Genta (June 2011). "Pectínidos de la Formación Paraná (Mioceno) Entre Ríos, Argentina". Serie Correlación Geológica (in Spanish). 27 (1): 66–75.
Péreza, Leandro M.; Iturrería, Santiago F. Genta; Griffini, Miguel (October 2010). "Paleoecological and Paleobiogeographic Significance of Two New Species of Bivalves in the Paraná Formation (Late Miocene) of Entre Ríos Province, Argentina". Malacologia. 53 (1): 61–76. doi:10.4002/040.053.0104. ISSN 0076-2997. S2CID 85017087.
Muravchik, Martín; Griffin, Miguel (2004). "Bryozoans from the Paraná Formation (Miocene), in Entre Ríos province, Argentina" (PDF). Ameghiniana (in Spanish). 41 (1): 3–12. ISSN 1851-8044.
Martínez, Sergio; Pérez, Leandro M.; del Río, Claudia J. (2021-10-02). "Echinoderms from the upper Miocene Paraná Formation of Argentina". Alcheringa: An Australasian Journal of Palaeontology. 45 (4): 435–442. doi:10.1080/03115518.2021.1985609. ISSN 0311-5518. S2CID 243479952.
Franco, María Jimena (2008-11-30). "Leños fósiles de la Formación Paraná (Mioceno Medio), Toma Vieja, Paraná, Entre Ríos, Argentina: registro de bosques estacionales mixtos". Ameghiniana. 45 (4): 669–717.
Anzótegui, Luisa M.; Aceñolaza, Pablo G. (2008-05-14). "Macrofloristic assemblage of the Parana Formation (Middle-Upper Miocene) in Entre Rios (Argentina)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 248 (2): 159–170. doi:10.1127/0077-7749/2008/0248-0159. hdl:11336/80848. ISSN 0077-7749.
Aceñolaza, Pablo G.; Aceñolaza, Florencio G. (1996). "IMPRONTAS FOLIARES DE UNA LAURACEAE EN LA FORMACIÓN PARANÁ (MIOCENO SUPERIOR), EN VILLA URQUIZA, ENTRE RÍOS" (in Spanish). 33 (2): 155–159. ISSN 1851-8044. {{cite journal}}: Cite journal requires |journal= (help)
Franco, M. Jimena; Brea, Mariana (2023-02-27). "Redescription of Solanumxylon paranensis, Late Miocene of Paraná Formation (Entre Ríos, Argentina): A Reliable Fossil Wood of Solanaceae". Ameghiniana. 60 (1). doi:10.5710/AMGH.13.10.2022.3518. ISSN 0002-7014. S2CID 253105168.

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[:17-1] Agnolin, Federico; Lucero, Sergio (December 2004). "Registros de cetáceos del Mioceno Tardío de Entre Ríos, Argentina". Revista de biología marina y oceanografía. 39 (2). doi:10.4067/S0718-19572004000200007. ISSN 0718-1957.

[:0-2] Pérez, Leandro M.; Cione, Alberto L.; Cozzuol, Mario; Varela, Augusto N. (June 2011). "A Sperm Whale (Cetacea: Physeteroidea) from the Paraná Formation (Late Miocene) of Entre Rís, Argentina. Environment and Taphonomy". Ameghiniana. 48 (4): 648–654. doi:10.5710/AMGH.v48i3(425). ISSN 0002-7014. S2CID 129118367.

[:9-3] Noriega, Jorge Ignacio; Cione, Alberto Luis; Aceñolaza, Florencio Gilberto (2007-09-05). "Shark tooth marks on Miocene balaenopterid cetacean bones from Argentina". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 245 (2): 185–192. doi:10.1127/0077-7749/2007/0245-0185. ISSN 0077-7749.

[4] Cozzuol, Mario A. (April 1996). "The record of aquatic mammals in southern South America". Münchner Geowiss. Abh. 30: 321–342.

[:10-5] Muizon, Christian de; Hendey, Q. B. (1980). "Late Tertiary seals of the south Atlantic Ocean". Annals of the South African Museum. Annale van die Suid-Afrikaanse Museum. 82: 91–128. ISSN 0303-2515.

[6] Muizon, Christian; Bond, Mariano (1982). "Le Phocidae (Mammalia) de la Formation Parana (Entre Rios, Argentine)". Bull. Mus. Natn. Hist. Nat., Paris (in French): 165–207.

[:1-7] Candela, Adriana M.; Bonini, Ricardo A.; Noriega, Jorge I. (November 2012). "First continental vertebrates from the marine Paraná Formation (Late Miocene, Mesopotamia, Argentina): Chronology, biogeography, and paleoenvironments". Geobios. 45 (6): 515–526. Bibcode:2012Geobi..45..515C. doi:10.1016/j.geobios.2012.05.003. S2CID 128431992.

[:3-8] Vélez-Juarbe, Jorge; Noriega, Jorge I.; Ferrero, Brenda S. (2012). "FOSSIL DUGONGIDAE (MAMMALIA, SIRENIA) FROM THE PARANÁ FORMATION (LATE MIOCENE) OF ENTRE RÍOS PROVINCE, ARGENTINA". Ameghiniana. 49 (4): 585–593. doi:10.5710/AMGH.27.6.2012.568. hdl:11336/72358. ISSN 0002-7014. S2CID 56435219.

[:2-9] Noriega, Jorge I.; Cenizo, Marcos; Brandoni, Diego; Pérez, Leandro M.; Tineo, David E.; Diederle, Juan M.; Bona, Paula (2023-05-09). "A new pelican (Aves: Pelecanidae) from the Upper Miocene of Argentina: new clues about the origin of the New World lineages". Journal of Vertebrate Paleontology. doi:10.1080/02724634.2023.2202702. ISSN 0272-4634. S2CID 258605013.

[:6-10] Woodward, Arthur Smith (1900). "I.—On some fish-remains from the Parana formation, Argentine Republic". The Annals and Magazine of Natural History; Zoology, Botany, and Geology. 6 (31): 1–7. doi:10.1080/00222930008678334. ISSN 0374-5481.

[:16-11] "Tetraodontiformes (Osteichthyes, Actinopterygii) from the Miocene of Argentina: with the southernmost record of fossil Tetraodontidae". Comptes Rendus Palevol. 20 (27): 585–596. 2021.

[12] "Squalus obliquidens | Shark-References". shark-references.com. Retrieved 2023-05-21.

[:5-13] Cione, Alberto Luis; Cabrera, Daniel Alfredo; Barla, María Julia (2012-03-01). "Oldest record of the Great White Shark (Lamnidae, Carcharodon; Miocene) in the Southern Atlantic". Geobios. 45 (2): 167–172. Bibcode:2012Geobi..45..167C. doi:10.1016/j.geobios.2011.06.002. ISSN 0016-6995.

[:4-14] Cione, Alberto Luis; Mennucci, Jorge; P&eacute, Leandro; rez; Marí Barla, a Julia (2008-01-01). "Megascyliorhinus trelewensis (Neoselachii) in the Middle-Upper Miocene of Parana, Central Eastern Argentina". Instituto Superior de Correlacion Geologica, Serie Miscelanea. 17 (1): 41–49.

[:13-15] Martín Pérez, Leandro; Griffin, Miguel; Iturerría, Santiago Genta (June 2011). "Pectínidos de la Formación Paraná (Mioceno) Entre Ríos, Argentina". Serie Correlación Geológica (in Spanish). 27 (1): 66–75.

[:12-16] Péreza, Leandro M.; Iturrería, Santiago F. Genta; Griffini, Miguel (October 2010). "Paleoecological and Paleobiogeographic Significance of Two New Species of Bivalves in the Paraná Formation (Late Miocene) of Entre Ríos Province, Argentina". Malacologia. 53 (1): 61–76. doi:10.4002/040.053.0104. ISSN 0076-2997. S2CID 85017087.

[:15-17] Muravchik, Martín; Griffin, Miguel (2004). "Bryozoans from the Paraná Formation (Miocene), in Entre Ríos province, Argentina" (PDF). Ameghiniana (in Spanish). 41 (1): 3–12. ISSN 1851-8044.

[:14-18] Martínez, Sergio; Pérez, Leandro M.; del Río, Claudia J. (2021-10-02). "Echinoderms from the upper Miocene Paraná Formation of Argentina". Alcheringa: An Australasian Journal of Palaeontology. 45 (4): 435–442. doi:10.1080/03115518.2021.1985609. ISSN 0311-5518. S2CID 243479952.

[:7-19] Franco, María Jimena (2008-11-30). "Leños fósiles de la Formación Paraná (Mioceno Medio), Toma Vieja, Paraná, Entre Ríos, Argentina: registro de bosques estacionales mixtos". Ameghiniana. 45 (4): 669–717.

[:11-20] Anzótegui, Luisa M.; Aceñolaza, Pablo G. (2008-05-14). "Macrofloristic assemblage of the Parana Formation (Middle-Upper Miocene) in Entre Rios (Argentina)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 248 (2): 159–170. doi:10.1127/0077-7749/2008/0248-0159. hdl:11336/80848. ISSN 0077-7749.

[:8-21] Aceñolaza, Pablo G.; Aceñolaza, Florencio G. (1996). "IMPRONTAS FOLIARES DE UNA LAURACEAE EN LA FORMACIÓN PARANÁ (MIOCENO SUPERIOR), EN VILLA URQUIZA, ENTRE RÍOS" (in Spanish). 33 (2): 155–159. ISSN 1851-8044. {{cite journal}}: Cite journal requires |journal= (help)

[22] Franco, M. Jimena; Brea, Mariana (2023-02-27). "Redescription of Solanumxylon paranensis, Late Miocene of Paraná Formation (Entre Ríos, Argentina): A Reliable Fossil Wood of Solanaceae". Ameghiniana. 60 (1). doi:10.5710/AMGH.13.10.2022.3518. ISSN 0002-7014. S2CID 253105168.