Platytholus

Platytholus; Temporal range: Late Cretaceous,
	Life restoration
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	†Ornithischia
Suborder:	†Pachycephalosauria
Family:	†Pachycephalosauridae
Subfamily:	†Pachycephalosaurinae
Genus:	†Platytholus; Horner, Goodwin & Evans, 2023
Species:	†P. clemensi
Binomial name
	†Platytholus clemensi; Horner, Goodwin & Evans, 2023

Platytholus is an extinct genus of pachycephalosaurid dinosaur from the Late Cretaceous (Maastrichtian) Hell Creek Formation of the United States. The genus contains a single species, P. clemensi, known from a partial skull.[1]

Discovery and naming

The Platytholus holotype specimen, MOR 2915, was discovered during the early 2000s in sediments of the Hell Creek Formation, dated to the latest Maastrichtian age of the late Cretaceous period, in Garfield County, Montana. The specimen consists of a partial skull.[1]

In 2023, Horner, Goodwin & Evans described Platytholus clemensi, a new genus and species of pachycephalosaurid, based on these fossil remains. The generic name, "Platytholus", combines the Latinised Greek "platys", meaning "flattened", "wide", or "broad", and "tholos", meaning a "small domed hill". The specific name, "clemensi", honors American paleontologist William A. Clemens Jr.[1]

Classification

Horner, Goodwin & Evans (2023) recovered Platytholus as a pachycephalosaurine member of the Pachycephalosauridae, as the sister taxon to a clade formed by Prenocephale and Acrotholus. The results of their phylogenetic analyses are shown in the cladogram below:[1]

Pachycephalosauria

Wannanosaurus

Pachycephalosauridae

	Hanssuesia

	Colepiocephale

	Stegoceras

Pachycephalosaurinae

Goyocephale

Homalocephale

Tylocephale

Foraminacephale

Amtocephale

	Prenocephale

	Acrotholus

	Platytholus

	Sphaerotholus edmontonensis

	Sphaerotholus goodwini

	Sphaerotholus buchholtzae

	Pachycephalosaurus

	Alaskacephale

References

Horner, John R.; Goodwin, Mark B.; Evans, David C. (2023-04-14). "A new pachycephalosaurid from the Hell Creek Formation, Garfield County, Montana, U.S.A.". Journal of Vertebrate Paleontology: e2190369. doi:10.1080/02724634.2023.2190369. ISSN 0272-4634.

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[Platytholus-1] Horner, John R.; Goodwin, Mark B.; Evans, David C. (2023-04-14). "A new pachycephalosaurid from the Hell Creek Formation, Garfield County, Montana, U.S.A.". Journal of Vertebrate Paleontology: e2190369. doi:10.1080/02724634.2023.2190369. ISSN 0272-4634.