Pomatiopsidae

Species in the family Pomatiopsidae occur worldwide.[1] The generic diversity of Pomatiopsinae is particularly high in the Japanese Archipelago, where four of the eight genera, including two endemics, are recorded.[1] The subfamily Triculinae radiated as aquatic snails in freshwater habitats in Southeast Asia.[1]

Approximate distribution map of Pomatiopsidae.
Notes: Distribution of Tomichia includes also Central Africa,[2]
Cecina has eight species.[3]

The American malacologist William Stimpson first defined this taxon as Pomatiopsinae in 1865.[4] Stimpson's diagnosis reads as follows:[4]

Pomatiopsinae, with the shell and operculum as in the Rissoinae. Foot with lateral sinuses. Size small. Amphibious. Genus Pomatiopsis, Tryon.

The family Pomatiopsidae consists of 2 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005)[5] that follows classification by Davis (1979):[6]

• Subfamily Pomatiopsinae Stimpson, 1865 - synonyms: Hemibiinae Heude, 1890; Tomichiinae Wenz, 1938;[7] Coxiellidae Iredale, 1943;[8] Oncomelaniidae Salisbury & Edwards, 1961; Cecininae Starobogatov, 1983
• Subfamily Triculinae Annandale, 1924[9]
  • tribe Triculuni Annandale, 1924 - synonym: Delavayidae Annandale, 1924[10]
  • tribe Jullieniini Davis, 1979[6]
  • tribe Lacunopsini Davis, 1979[6]
  • tribe Pachydrobiini Davis & Kang, 1990[11]

Family-group name Rehderiellinae Brandt, 1974[12] is also in Pomatiopsidae, but it is not allocated in detail.[5]

Genera within the family Pomatiopsidae include:

• Spiripockia Simone, 2012 (not allocated to a subfamily)[13]

Subfamily Pomatiopsinae

• Blanfordia Adams, 1863[1]
• Cecina A. Adams, 1861[5][14]
• Coxiella E. A. Smith, 1894:[5] belongs in the family Tomichiidae
• Floridiscrobs Pilsbry and McGinty, 1949
• Fukuia Abbott & Hunter, 1949[1]
• Hemibia Heude, 1890[5] (species type: Oncomelania hupensis Gredler, 1881
• Idiopyrgus Pilsbry, 1911[1][15] - synonym: Aquidauania Davis, 1979:[6] belongs in the family Tomichiidae
• Oncomelania Gredler, 1881[5]
• Pomatiopsis Tryon, 1862 - the type genus of the family Pomatiopsidae[5]
• Tomichia Benson, 1851:[5][16] belongs in the family Tomichiidae

Subfamily Triculinae - there are over 20 genera in Triculinae[1]

tribe Triculuni

• Delavaya Heude, 1889[5][11]
• Fenouilia Heude, 1889[11]
• Lithoglyphopsis Thiele, 1928[17][18]
• Tricula Benson, 1843 - type genus of the tribe Triculuni[5][11]

tribe Jullieniini

• Hubendickia Brandt, 1968[6]
• Hydrorissoia Bavay, 1895[6]
• Jullienia Crosse & P. Fischer, 1876 - type genus of the tribe Jullieniini[5]
• Karelainia Davis, 1979[6]
• Kunmingia Davis & Kuo in Davis, 1981[17][18]
• Neoprososthenia Davis & Kuo in Davis, Kuo, Hoagland, Chen, Yang & Chen, 1984[17][18]
• Pachydrobiella Thiele, 1928[6] - with the only species Pachydrobiella brevis (Bavay, 1895)[6]
• † Paraprososthenia Annandale, 1919[6]
• Saduniella Brandt, 1970[6] - with the only species Saduniella planispira Brandt, 1970[6]

tribe Lacunopsini

• Lacunopsis Deshayes, 1876 - type genus of the tribe Lacunopsini[5]

tribe Pachydrobiini

• Gammatricula Davis & Liu in Davis, Liu & Chen, 1990[19]
• Halewisia Davis, 1979[11] - with the only species Halewisia expansa (Brandt, 1970)[6]
• Jinghongia Davis in Davis & Kang, 1990[11]
• Neotricula Davis in Davis, Subba Rao & Hoagland, 1986[11]
• Pachydrobia Crosse & P. Fischer, 1876[11] - type genus of the tribe Pachydrobiini[5]
• Robertsiella Davis & Greer, 1980[11]
• Wuconchona Kang, 1983[11]

Rehderiellinae is not allocated to a subfamily[5]

• Rehderiella Brandt, 1974 - type genus of the taxon Rehderiellinae[5]

The Pomatiopsidae have various life habits: aquatic, amphibious, littoral, halophilic, cavernicolous and even terrestrial.[1][13] Terrestrial taxa occur only on the Japanese Archipelago located in East Asia (Blanfordia).[1] Tomichia and Coxiella include several halophilic species occurring on saline lakes.[1]

Pomatiopsidae invaded freshwater habitats from marine ones in one or in two independent lineages.[20] They also invaded terrestrial habitats from freshwater habitats in two independent lineages.[1]

This article incorporates public domain text from the reference[4] and CC-BY-2.0 from the reference[1]

Wikimedia Commons has media related to Pomatiopsidae.

• Stimpson W. (1865). "Researches upon the Hydrobiinae and allied forms chiefly made upon materials in the museum of the Smithsonian Institution". Smithsonian Miscellaneous Collections 7(201): 1-59. page 4
• Davis G. M., Chen C.-E., Wu C., Kuang T.-F., Xing X.-G., Li L., Liu W.-J. & Yan Y.-L. (1992). "The Pomatiopsidae of Hunan, China (Gastropoda, Rissoacea)". Malacologia 34(1-2): 143-342.
• Guan, F.; Niu, A. O.; Attwood, S. W.; Li, Y. L.; Zhang, B.; Zhu, Y. H. (2008). "Molecular phylogenetics of Triculine snails (Gastropoda: Pomatiopsidae) from southern China". Molecular Phylogenetics and Evolution. 48 (2): 702–707. doi:10.1016/j.ympev.2008.04.021. PMID 18502667.
• Liu, L.; Huo, G.-N.; He, H.-B.; Zhou, B.; Attwood, S. W. (2014). "A phylogeny for the Pomatiopsidae (Gastropoda: Rissooidea): a resource for taxonomic, parasitological and biodiversity studies". BMC Evolutionary Biology. 14 (1): 29. doi:10.1186/1471-2148-14-29. PMC 4016560. PMID 24548800.