Trimma

Trimma
	Trimma gigantum
	Trimma caesiura
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Gobiiformes
Family:	Gobiidae
Genus:	Trimma; D. S. Jordan & Seale, 1906
Type species
	Trimma caesiura; D. S. Jordan & Seale, 1906

Trimma is a genus of fish in the family Gobiidae native to the Indian and Pacific Ocean. Together with members of the genus Eviota, they are known commonly as pygmygobies or dwarfgobies.

Species

Trimma caudomaculatum

Trimma habrum

Trimma lantana

Trimma nasa

Trimma naudei female 23.3 mm, Nha Trang, Vietnam

Trimma okinawae

Trimma tevegae

There are currently 98 recognized species in this genus:

Trimma abyssum G. R. Allen, 2015 (Deep-reef pygmygoby) [1]
Trimma agrena R. Winterbottom & I. S. Chen, 2004 (Fishnet pygmygoby)
Trimma anaima R. Winterbottom, 2000 (Sharp-eye pygmygoby) [2]
Trimma annosum R. Winterbottom, 2003 (Grey-bearded pygmygoby) [2]
Trimma anthrenum R. Winterbottom, 2006 (Honey-bee pygmygoby)
Trimma aturirii R. Winterbottom, Erdmann & Cahyani, 2015 (Aturiri's pygmygoby) [3]
Trimma avidori (Goren, 1978)
Trimma barralli R. Winterbottom, 1995
Trimma benjamini R. Winterbottom, 1996 (Ring-eye pygmygoby) [2]
Trimma bisella R. Winterbottom, 2000
Trimma burridgeae R. Winterbottom, 2016 [4]
Trimma caesiura D. S. Jordan & Seale, 1906 (Caesiura pygmygoby) [2]
Trimma cana R. Winterbottom, 2004 (Candycane pygmygoby)
Trimma capostriatum (Goren, 1981) (Spotted red-lined pygmygoby) [2]
Trimma caudipunctatum T. Suzuki & Senou, 2009
Trimma caudomaculatum Yoshino & Araga, 1975 [4]
Trimma cheni R. Winterbottom, 2011 (Face-stripe pygmygoby)
Trimma chledophilum G. R. Allen, 2015 (Mud pygmygoby) [1]
Trimma corallinum (J. L. B. Smith, 1959) (Polka-dot pygmygoby)
Trimma corerefum R. Winterbottom, 2016 [4]
Trimma dalerocheila R. Winterbottom, 1984
Trimma emeryi R. Winterbottom, 1985 (Emery's pygmygoby) [2]
Trimma erdmanni R. Winterbottom, 2011 (Erdmann's pygmygoby)
Trimma erwani Viviani, J. T. Williams & Planes, 2016 (Slanted pygmygoby) [5]
Trimma fangi R. Winterbottom & I. S. Chen, 2004 (Spot-faced pygmygoby) [2]
Trimma fasciatum T. Suzuki, Sakaue & Senou, 2012 [6]
Trimma filamentosus R. Winterbottom, 1995
Trimma fishelsoni Goren, 1985
Trimma flammeum (J. L. B. Smith, 1959)
Trimma flavatrum K. Hagiwara & R. Winterbottom, 2007 (Wasp pygmygoby) [2]
Trimma flavicaudatum (Goren, 1982)
Trimma fraena R. Winterbottom, 1984
Trimma fucatum R. Winterbottom & Southcott, 2007 (Harlot pygmygoby)
Trimma gigantum R. Winterbottom & Zur, 2007 (Giant pygmygoby)
Trimma grammistes (Tomiyama, 1936)
Trimma griffithsi R. Winterbottom, 1984 (Griffiths' pygmygoby)
Trimma habrum R. Winterbottom, 2011 (Delicate pygmygoby)
Trimma haima R. Winterbottom, 1984
Trimma haimassum R. Winterbottom, 2011 (Blood-spot pygmygoby)
Trimma halonevum R. Winterbottom, 2000 (Red-spot pygmygoby)
Trimma hayashii K. Hagiwara & R. Winterbottom, 2007 (Four-eye pygmygoby)
Trimma helenae R. Winterbottom, Erdmann & Cahyani, 2014 (Helen's pygmygoby) [7]
Trimma hoesei R. Winterbottom, 1984 (Fork-tail pygmygoby) [2]
Trimma hollemani R. Winterbottom, 2016 [4]
Trimma hotsarihiensis R. Winterbottom, 2009 (Helen Reef pygmygoby)
Trimma imaii T. Suzuki & Senou, 2009 (Imai's pygmygoby)
Trimma insularum R. Winterbottom & Hoese, 2015 (Cocos pygmygoby) [2]
Trimma irinae R. Winterbottom, 2014 (Irina's pygmygoby) [8]
Trimma kardium R. Winterbottom, Erdmann & Cahyani, 2015 (Heart pygmygoby) [3]
Trimma kitrinum R. Winterbottom & Hoese, 2015 (Citron pygmygoby) [2]
Trimma kudoi T. Suzuki & Senou, 2008
Trimma lantana R. Winterbottom & C. A. Villa, 2003 (Lantana pygmygoby) [2]
Trimma lutea Viviani, J. T. Williams & Planes, 2016 (Yellow-barred pygmygoby) [5]
Trimma macrophthalmus (Tomiyama, 1936) (Large-eye pygmygoby) [2]
Trimma maiandros Hoese, R. Winterbottom & Reader, 2011 (Zigzag pygmygoby) [2]
Trimma marinae R. Winterbottom, 2005 (Princess pygmygoby)
Trimma matsunoi T. Suzuki, Sakaue & Senou, 2012 [6]
Trimma mendelssohni (Goren, 1978)
Trimma meranyx R. Winterbottom, Erdmann & Cahyani, 2014 (Day-night pygmygoby) [9]
Trimma meristum R. Winterbottom & Hoese, 2015 (Split-ray pygmygoby) [2]
Trimma milta R. Winterbottom, 2002 (Red-earth pygmygoby) [2]
Trimma multiclitellum G. R. Allen, 2015 (Multi-saddle pygmygoby) [1]
Trimma nasa R. Winterbottom, 2005 (Nasal-bar pygmygoby) [2]
Trimma nauagium G. R. Allen, 2015 (Shipwreck pygmygoby) [1]
Trimma naudei J. L. B. Smith, 1957 (Naude's pygmygoby)
Trimma necopinum (Whitley, 1959) (Australian pygmygoby) [2]
Trimma nomurai T. Suzuki & Senou, 2007 (Lilac pygmygoby) [2]
Trimma okinawae (Aoyagi, 1949) (Orange-red pygmygoby) [2]
Trimma omanensis R. Winterbottom, 2000
Trimma pajama R. Winterbottom, Erdmann & Cahyani, 2014 (Pajama pygmygoby) [9]
Trimma panemorfum R. Winterbottom & Pyle, 2004[10]
Trimma papayum R. Winterbottom, 2011 (Pawpaw pygmygoby)
Trimma pentherum R. Winterbottom & Hoese, 2015 (Mourning pygmygoby) [2]
Trimma preclarum R. Winterbottom, 2006 (Exquisite pygmygoby) [2]
Trimma quadrimaculatum Hoese, Bogorodsky & A. O. Mal, 2015 (Four-spotted pygmygoby) [11]
Trimma randalli R. Winterbottom & Zur, 2007 (Randall's pygmygoby)
Trimma readerae R. Winterbottom & Hoese, 2015 [2]
Trimma rubromaculatum G. R. Allen & Munday, 1995 (Red-spotted dwarfgoby)
Trimma sanguinellus R. Winterbottom & Southcott, 2007 (Sanguinello pygmygoby)
Trimma sheppardi R. Winterbottom, 1984 (Sheppard's pygmygoby)
Trimma sostra R. Winterbottom, 2004 (Sostra pygmygoby)
Trimma squamicana R. Winterbottom, 2004 (Candy-scale pygmygoby)
Trimma stobbsi R. Winterbottom, 2001 (Stobb's pygmygoby) [2]
Trimma striatum (Herre, 1945) (Red-lined pygmygoby) [2]
Trimma tauroculum R. Winterbottom & Zur, 2007 (Bulls-eye pygmygoby)
Trimma taylori Lobel, 1979 (Yellow pygmygoby) [2]
Trimma tevegae Cohen & W. P. Davis, 1969 (Blue-stripe pygmygoby) [4]
Trimma trioculatum R. Winterbottom, Erdmann & Cahyani, 2015 (Three-eyed pygmygoby) [3]
Trimma unisquame (Gosline, 1959) (Black-margin pygmygoby) [2]
Trimma volcana R. Winterbottom, 2003 (Volcano pygmygoby)
Trimma winchi R. Winterbottom, 1984
Trimma winterbottomi J. E. Randall & Downing, 1994 (Winterbottom's pygmygoby)
Trimma woutsi R. Winterbottom, 2002
Trimma xanthochrum R. Winterbottom, 2011 (Step-spot pygmygoby)
Trimma xanthum R. Winterbottom & Hoese, 2015 (Yellow-red pygmygoby) [2]
Trimma yanagitai T. Suzuki & Senou, 2007
Trimma yanoi T. Suzuki & Senou, 2008 (Yano's pygmygoby)
Trimma yoshinoi T. Suzuki, Ko. Yano & Senou, 2015 [12]
Trimma zurae R. Winterbottom, Erdmann & Cahyani, 2014 (Marg's pygmygoby) [9]

References

Allen, G.R. (2015): Four new species of Trimma (Pisces: Gobiidae) from Papua New Guinea and Indonesia. Journal of the Ocean Science Foundation, 14: 28-48.
Winterbottom, R. & Hoese, D.F. (2015): A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species. Zootaxa, 3934 (1): 1–102.
Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2015): New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature. Zootaxa, 3973 (2): 201–226.
Winterbottom, R. (2016): Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa, 4144 (1): 1-53.
Viviani, J., Williams, J.T. & Planes, S. (2016): Two new pygmygobies (Percomorpha: Gobiidae: Trimma) from French Polynesia. Journal of the Ocean Science Foundation, 23: 1–11.
Suzuki, T., Sakaue, J. & Senou, H. (2012): Two New Species of the Gobiid Fish Genus Trimma (Actinopterygii: Perciformes: Gobiidae) from Japan and Palau. Bulletin of the National Museum of Nature and Science, Ser. A, Supplement No. 6: 67-77.
Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2014): Trimma helenae (Pisces; Gobioidei), a new species of gobiid fish from Indonesia. Zootaxa, 3760 (3): 420–428.
Winterbottom, R. (2014): Trimma irinae, (Pisces; Gobioidei) a new species of gobiid fish from Milne Bay Province, Papua New Guinea. Zootaxa, 3802 (2): 209–216.
Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2014): Three new species of Trimma (Pisces; Gobioidei) from Indonesia. Zootaxa, 3838 (3): 367–384.
WINTERBOTTOM, R. ., & PYLE, R. L. . (2022). A new species of Trimma (Pisces: Gobiidae) from the deep reefs of Palau, western Pacific Ocean. Zootaxa, 5094(4), 595–600. https://doi.org/10.11646/zootaxa.5094.4.5
Hoese, D.F., Bogorodsky, S.V. & Mal, A.O. (2015): Description of a new species of Trimma (Perciformes: Gobiidae) from the Red Sea, with a discussion of the generic separation of Trimma and Priolepis, with discussion of sensory papillae terminology. Zootaxa, 4027 (4): 538–550.
Suzuki, T., Yano, K. & Senou, H. (2015): Trimma yoshinoi, a new gobiid fish from Japan (Perciformes: Gobiidae). Journal of the Ocean Science Foundation, 14: 66-73.

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[Allen2015-1] Allen, G.R. (2015): Four new species of Trimma (Pisces: Gobiidae) from Papua New Guinea and Indonesia. Journal of the Ocean Science Foundation, 14: 28-48.

[Winterbottom2015-2] Winterbottom, R. & Hoese, D.F. (2015): A revision of the Australian species of Trimma (Actinopterygii, Gobiidae), with descriptions of six new species and redescriptions of twenty-three valid species. Zootaxa, 3934 (1): 1–102.

[Winterbottom2015a-3] Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2015): New species of Trimma (Actinopterygii, Gobiidae) from Indonesia, with comments on head papillae nomenclature. Zootaxa, 3973 (2): 201–226.

[Winterbottom2016-4] Winterbottom, R. (2016): Trimma tevegae and T. caudomaculatum revisited and redescribed (Acanthopterygii, Gobiidae), with descriptions of three new similar species from the western Pacific. Zootaxa, 4144 (1): 1-53.

[Viviani2016-5] Viviani, J., Williams, J.T. & Planes, S. (2016): Two new pygmygobies (Percomorpha: Gobiidae: Trimma) from French Polynesia. Journal of the Ocean Science Foundation, 23: 1–11.

[Suzuki2012-6] Suzuki, T., Sakaue, J. & Senou, H. (2012): Two New Species of the Gobiid Fish Genus Trimma (Actinopterygii: Perciformes: Gobiidae) from Japan and Palau. Bulletin of the National Museum of Nature and Science, Ser. A, Supplement No. 6: 67-77.

[Winterbottom2014-7] Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2014): Trimma helenae (Pisces; Gobioidei), a new species of gobiid fish from Indonesia. Zootaxa, 3760 (3): 420–428.

[Winterbottom2014a-8] Winterbottom, R. (2014): Trimma irinae, (Pisces; Gobioidei) a new species of gobiid fish from Milne Bay Province, Papua New Guinea. Zootaxa, 3802 (2): 209–216.

[Winterbottom2014b-9] Winterbottom, R., Erdmann, M.V. & Dita Cahyani, N.K. (2014): Three new species of Trimma (Pisces; Gobioidei) from Indonesia. Zootaxa, 3838 (3): 367–384.

[10] WINTERBOTTOM, R. ., & PYLE, R. L. . (2022). A new species of Trimma (Pisces: Gobiidae) from the deep reefs of Palau, western Pacific Ocean. Zootaxa, 5094(4), 595–600. https://doi.org/10.11646/zootaxa.5094.4.5

[Hoese2015-11] Hoese, D.F., Bogorodsky, S.V. & Mal, A.O. (2015): Description of a new species of Trimma (Perciformes: Gobiidae) from the Red Sea, with a discussion of the generic separation of Trimma and Priolepis, with discussion of sensory papillae terminology. Zootaxa, 4027 (4): 538–550.

[Suzuki2015-12] Suzuki, T., Yano, K. & Senou, H. (2015): Trimma yoshinoi, a new gobiid fish from Japan (Perciformes: Gobiidae). Journal of the Ocean Science Foundation, 14: 66-73.