Trionychia

Trionychia is a superfamily of turtles which encompasses the species that are commonly referred to as softshelled turtles as well as some others. The group contains two families, Carettochelyidae, which has only one living species, the pig-nosed turtle (Carettochelys insculpta) native to New Guinea and Northern Australia, and Trionychidae, the softshelled turtles, containing numerous species native to Asia, North America and Africa. These families likely diverged during the late Jurassic.[2] The oldest known stem-trionychian is Sinaspideretes from the Late Jurassic of China.[3][4]

Trionychia Temporal range: Late Jurassic to recent,

The spiny softshell turtle (Trionychidae)

The pig-nosed turtle (Carettochelyidae)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade:	Polycryptodira
Superfamily:	Trionychia Hummel, 1929[1]
Families
Carettochelyidae Trionychidae and see text
Synonyms
Trionychoidea

Systematics

Except for those not assigned to a family, only living genera are listed here.

Family Carettochelyidae
- Subfamily Carettochelyinae
  - Genus Carettochelys
Pan-Trionychidae
- Family Plastomenidae[5] (fossil)
- Family Trionychidae
  - Subfamily Cyclanorbinae
    - Genus Cyclanorbis
    - Genus Cycloderma
    - Genus Lissemys
  - Subfamily Trionychinae
    - Genus Amyda
    - Genus Apalone
    - Genus Chitra
    - Genus Dogania
    - Genus Nilssonia
    - Genus Palea
    - Genus Pelochelys
    - Genus Pelodiscus
    - Genus Rafetus
    - Genus Trionyx

The eastern mud turtle (Kinosternon subrubrum) is a primitive relative of the true softshells which still has a typical hard turtle shell

References

Rhodin 2011, p. 000.204
Pereira, Anieli G.; Sterli, Juliana; Moreira, Filipe R.R.; Schrago, Carlos G. (August 2017). "Multilocus phylogeny and statistical biogeography clarify the evolutionary history of major lineages of turtles". Molecular Phylogenetics and Evolution. 113: 59–66. doi:10.1016/j.ympev.2017.05.008. ISSN 1055-7903. PMID 28501611.
Ouyang, Hui; Li, Lu; Tong, Haiyan (July 2014). "A revision of Sinaspideretes wimani Young & Chow, 1953 (Testudines: Cryptodira: Trionychoidae) from the Jurassic of the Sichuan Basin, China". Geological Magazine. 151 (4): 600–610. Bibcode:2014GeoM..151..600T. doi:10.1017/S0016756813000575. ISSN 0016-7568. S2CID 128423062.
Evers, Serjoscha W.; Benson, Roger B. J. (January 2019). Smith, Andrew (ed.). "A new phylogenetic hypothesis of turtles with implications for the timing and number of evolutionary transitions to marine lifestyles in the group". Palaeontology. 62 (1): 93–134. doi:10.1111/pala.12384. S2CID 134736808.
One or more basal lineages formerly believed to be a distinct family

Bibliography

Rhodin, Anders G.J.; van Dijk, Peter Paul; Inverson, John B.; Shaffer, H. Bradley; Roger, Bour (2011-12-31). "Turtles of the world, 2011 update: Annotated checklist of taxonomy, synonymy, distribution and conservation status" (PDF). Chelonian Research Monographs. 5. Archived from the original (PDF) on 2012-01-31.

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Bouliachelys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Alienochelys †Berruchelus †Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Tortoises	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

†Baenoidea

†Baenidae	†Arvinachelys †Baena †Cedrobaena †Chisternon †Denazinemys †Gamerabaena †Neurankylus †Palatobaena †Peckemys †Plesiobaena
†Pleurosternidae	†Dinochelys †Dorsetochelys †Glyptops †Pleurosternon †Selenemys

Chelydridae	†Acherontemys Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Eurysternidae	†Achelonia †Chelonides †Eurysternum †Hydropelta †Idiochelys †Palaeomedusa †Parachelys
†Macrobaenidae	†Anatolemys †Aurorachelys †Kirgizemys †Osteopygis †Yakemys
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Archelon †Atlantochelys †Atlantochelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
†Sinemydidae	†Jeholochelys †Liaochelys †Manchurochelys †Ordosemys †Sinemys
†Thalassemydidae	†Neusticemys †Palaeomedusa †Thalassemys
	†Adocus †Angolachelys †Argillochelys †Bashuchelys †Brodiechelys †Ctenochelys †Compsemys †Hoyasemys †Larachelus †Leyvachelys †Plesiochelys †Prionochelys †Toxochelys †Uluops †Xinjiangchelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Chubutemys †Hispaniachelys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

Taxon identifiers	Wikidata: Q1877590 Wikispecies: Trionychoidea BioLib: 60940 CoL: 87BWZ Fauna Europaea: 15551 Fauna Europaea (new): 15bcf7d3-5f9f-4b86-8414-2b058e82b796 Fossilworks: 37646 ITIS: 948949

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