Carettochelyidae

Carettochelyidae; Temporal range:
	Pig-nosed turtle (Carettochelys insculpta)
	Fossil of Allaeochelys crassesculptata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Superfamily:	Trionychia
Family:	Carettochelyidae; Gill, 1889
Genera
	See text

Carettochelyidae is a family of cryptodiran turtles belonging to the Trionychia. It contains only a single living species, the pig-nosed turtle (Carettochelys insculpta) native to New Guinea and Northern Australia. Stem-group carettochelyids are known from the Cretaceous of Asia, with the family being widely distributed across North America, Europe and Asia during the Paleogene.

Taxonomy

After Joyce, 2014[1] unless otherwise noted.

Stem group taxa (also known as Pan-Carettochelys Joyce, Parham and Gauthier 2004)
Kizylkumemys Nessov, 1976
- Kizylkumemys khoratensis Tong et al., 2005 Khok Kruat Formation, Thailand, Early Cretaceous (Aptian)
- Kizylkumemys schultzi Nessov, 1976 Khodzhakul Formation, Uzbekistan, Late Cretaceous (Cenomanian)
Indeterminate fragments of stem-Carretochelyids are also known from the Cenomanian aged Bayan Shireh Formation, Mongolia
Carettochelyidae Gill, 1889
- Anosteira Leidy, 1871
  - Anosteira manchuriana Zangerl, 1947 Liaoning, China, late Eocene
  - Anosteira maomingensis Chow and Liu, 1955 Youkanwo Formation, Guangdong, China, Late Eocene
  - Anosteira mongoliensis Gilmore, 1931 Inner Mongolia, China, Late Eocene-Oligocene
  - Anosteira ornata Leidy, 1871 Bridger Formation, Wyoming, USA, Early Eocene
  - Anosteira pulchra (Clark, 1932) Uinta Formation, Utah, USA, Middle Eocene (Lutetian)
- An indeterminate carettochelyid is also known from the Early Eocene (Lutetian) aged Kuldana Formation, Khyber Pakhtunkhwa, Pakistan given the name Chorlakkichelys shahi Broin, 1987
- Carettochelyinae Williams, 1950
  - Allaeochelys Noulet, 1867
    - Allaeochelys crassesculpta (Harrassowitz, 1922) Messel Pit, Germany, Early Eocene
    - Allaeochelys delheidi (Dollo, 1886) Brussels Formation, Belgium, Zamora, Spain, Early Eocene Headon Hill Formation, England, Late Eocene (Priabonian)
    - Allaeochelys libyca Havlik et al, 2014, Al Wahat District, Libya, middle Miocene (Langhian)
    - Allaeochelys lingnanica (Young and Chow, 1962) Shaoguan, Guangdong, China early Paleogene (possibly Paleocene)
    - Allaeochelys magnifica (=Burmemys magnifica Hutchison et al., 2004) Pandaung Formation, Myanmar, Late Eocene (Bartonian)
    - Allaeochelys parayrei Noulet, 1867 Tarn, Toulouse, France, Late Eocene (Bartonian)
    - Allaeochelys liliae Carbot-Chanona et al. 2020[2] Mazantic Shale, Chiapas, Mexico, Early Miocene (Aquitanian)
  - Carettochelys Ramsay, 1886 Northern Australia, New Guinea, Recent
  - Indeterminate carettochelyids are also known from the uppermost Miocene to lowermost Pliocene of Victoria, Australia,[3] As well as the Upper Miocene of New Guinea.[4]

References

Joyce, Walter G. (April 2014). "A Review of the Fossil Record of Turtles of the Clade Pan-Carettochelys". Bulletin of the Peabody Museum of Natural History. 55 (1): 3–33. doi:10.3374/014.055.0102. ISSN 0079-032X. S2CID 59382889.
Carbot-Chanona, Gerardo; Rivera-Velázquez, Gustavo; Jiménez-Hidalgo, Eduardo; Reynoso, Víctor Hugo (December 2020). "The first Pan-Carettochelys turtle in the Neogene of the American continent and its paleobiogeographical relevance". Journal of South American Earth Sciences. 104: 102925. doi:10.1016/j.jsames.2020.102925. S2CID 224976641.
Rule, James P.; Kool, Lesley; Parker, William M.G.; Fitzgerald, Erich M.G. (2021-12-07). Mannion, Philip (ed.). "Turtles all the way down: Neogene pig‐nosed turtle fossil from southern Australia reveals cryptic freshwater turtle invasions and extinctions". Papers in Palaeontology. 8: spp2.1414. doi:10.1002/spp2.1414. ISSN 2056-2799. S2CID 245107305.
Glaessner, Martin F. (1942-07-08). "The occurrence of the New Guinea Turtle (Carettochelys) in the Miocene of Papua". Records of the Australian Museum. 21 (2): 106–109. doi:10.3853/j.0067-1975.21.1942.262. ISSN 0067-1975.

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[1] Joyce, Walter G. (April 2014). "A Review of the Fossil Record of Turtles of the Clade Pan-Carettochelys". Bulletin of the Peabody Museum of Natural History. 55 (1): 3–33. doi:10.3374/014.055.0102. ISSN 0079-032X. S2CID 59382889.

[2] Carbot-Chanona, Gerardo; Rivera-Velázquez, Gustavo; Jiménez-Hidalgo, Eduardo; Reynoso, Víctor Hugo (December 2020). "The first Pan-Carettochelys turtle in the Neogene of the American continent and its paleobiogeographical relevance". Journal of South American Earth Sciences. 104: 102925. doi:10.1016/j.jsames.2020.102925. S2CID 224976641.

[3] Rule, James P.; Kool, Lesley; Parker, William M.G.; Fitzgerald, Erich M.G. (2021-12-07). Mannion, Philip (ed.). "Turtles all the way down: Neogene pig‐nosed turtle fossil from southern Australia reveals cryptic freshwater turtle invasions and extinctions". Papers in Palaeontology. 8: spp2.1414. doi:10.1002/spp2.1414. ISSN 2056-2799. S2CID 245107305.

[4] Glaessner, Martin F. (1942-07-08). "The occurrence of the New Guinea Turtle (Carettochelys) in the Miocene of Papua". Records of the Australian Museum. 21 (2): 106–109. doi:10.3853/j.0067-1975.21.1942.262. ISSN 0067-1975.