Herichthys

Herichthys
	Herichthys cyanoguttatus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cichliformes
Family:	Cichlidae
Tribe:	Heroini
Genus:	Herichthys; S. F. Baird & Girard, 1854
Type species
	Herichthys cyanoguttatus; S. F. Baird & Girard, 1854

Herichthys is a small genus of cichlid fishes. Most are endemic to Mexico, but H. cyanoguttatus is also found in southern Texas (United States), and has been introduced to central Texas and Florida. In 2015, the genus was split, and 7 species moved into Nosferatu.

Species

There are currently 7 recognized species in this genus.[1] Separate mitochondrial DNA analyses by Husley et al.,[2] Rican et al.,[3] and Oldfield et al.,[4] support H. minckleyi as monophyletic and sister to all other Herichthys species. Moreover, molecular analysis conducted separately by López-Fernández, et al.,[5] and De la Maza-Benignos, et al.,[1] did not support H. tamasopoensis, H. cyanoguttatus, H. teporatus, and H. carpintis as distinct from each other. However, morphological and zoogeographical analysis by De la Maza-Benignos, et al.,[1] confirmed the validity of these four species, as well as differential evolutionary trends. Analysis also supported the separation of Herichthys into three distinct geomorphological groups + H. minckleyi. These are, from South to North:[1]

Herichthys deppii (Heckel, 1840) (Nautla cichlid)[1][6][7]
Herichthys tepehua De la Maza-Benignos, Ornelas-García, Lozano-Vilano, García-Ramírez & Doadrio, 2015[1][7]
Herichthys carpintis (D. S. Jordan & Snyder, 1899) (Pearlscale cichlid, lowland cichlid)
Herichthys tamasopoensis Artigas Azas, 1993 (Tamasopo cichlid)
Herichthys teporatus (Fowler, 1903) (Soto la Marina cichlid)[1][7]
Herichthys cyanoguttatus (S. F. Baird & Girard, 1854) (Texas cichlid, Rio Grande cichlid)
Herichthys minckleyi (Kornfield & J. N. Taylor, 1983) (Minkley's cichlid)

The Catalog of Fishes classifies the genus Nosferatu as synonymous with Herichthys.[8]

References

De la Maza-Benignos, M.; Ornelas-García, C.P.; Lozano-Vilano, M.d.L.; García-Ramírez, M.E. & Doadrio, I. (2015). "Phylogeographic analysis of genus Herichthys (Perciformes: Cichlidae), with descriptions of Nosferatu new genus and H. tepehua n. sp". Hydrobiologia. 748 (1): 201–231. doi:10.1007/s10750-014-1891-8. hdl:10261/126238. S2CID 16769534.
Hulsey, C.D., García, de León F.J., Sánchez-Jo Y. (2004). "Temporal diversification of Mesoamerican cichlid fishes across a major biogeographic boundary". Molecular Phylogenetics and Evolution. 31 (2): 754–764. doi:10.1016/j.ympev.2003.08.024. PMID 15062808.{{cite journal}}: CS1 maint: multiple names: authors list (link)
Rıcan O.; Pialek L.; Zardoya R.; Doadrio I.; Zrzavy J. (2013). "Biogeography of the Mesoamerican Cichlidae (Teleostei: Heroini): colonization through the GAARlandia land bridge and early diversification". Journal of Biogeography. 40 (3): 579–593. doi:10.1111/jbi.12023. hdl:10261/123988. S2CID 86520333.
Oldfield, R.G.; Mandrekar, K.; Nieves, M.X.; Hendrickson, D.A.; Chakrabarty, P.; Swanson, B.O.; Hofmann, H.A. (2015). "Parental care in the Cuatro Ciénegas cichlid, Herichthys minckleyi (Teleostei: Cichlidae)". Hydrobiologia. 748 (1): 233–257. doi:10.1007/s10750-014-2081-4. S2CID 17487888.
López-Fernández H.; Winemiller K.O.; Honeycutt R.L. (2010). "Multilocus phylogeny and rapid radiations in Neotropical cichlid fishes (Perciformes: Cichlidae)". Molecular Phylogenetics and Evolution. 55 (3): 1070–1086. doi:10.1016/j.ympev.2010.02.020. PMID 20178851.
Hans-Joachim P.; Morgenstern, R. & Schindler, I. (2014). "Cichlid fishes (Teleostei, Cichlidae) collected by Ferdinand Deppe in Mexico" (PDF). Vertebrate Zoology. 64 (1): 43–57. doi:10.3897/vz.64.e31462. S2CID 257187441. Archived from the original (PDF) on 2014-12-19. Retrieved 2014-12-19.
Mcmahan C. D.; Matamoros W. A.; Piller K. R.; Chakrabarty P. (2015). "Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera". Zootaxa. 3999 (2): 211–234. doi:10.11646/zootaxa.3999.2.3. PMID 26623572. S2CID 34699955.
Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Species in the genus Herichthys". Catalog of Fishes. California Academy of Sciences. Retrieved 10 November 2018.

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[MazaBenignos2015-1] De la Maza-Benignos, M.; Ornelas-García, C.P.; Lozano-Vilano, M.d.L.; García-Ramírez, M.E. & Doadrio, I. (2015). "Phylogeographic analysis of genus Herichthys (Perciformes: Cichlidae), with descriptions of Nosferatu new genus and H. tepehua n. sp". Hydrobiologia. 748 (1): 201–231. doi:10.1007/s10750-014-1891-8. hdl:10261/126238. S2CID 16769534.

[2] Hulsey, C.D., García, de León F.J., Sánchez-Jo Y. (2004). "Temporal diversification of Mesoamerican cichlid fishes across a major biogeographic boundary". Molecular Phylogenetics and Evolution. 31 (2): 754–764. doi:10.1016/j.ympev.2003.08.024. PMID 15062808.{{cite journal}}: CS1 maint: multiple names: authors list (link)

[3] Rıcan O.; Pialek L.; Zardoya R.; Doadrio I.; Zrzavy J. (2013). "Biogeography of the Mesoamerican Cichlidae (Teleostei: Heroini): colonization through the GAARlandia land bridge and early diversification". Journal of Biogeography. 40 (3): 579–593. doi:10.1111/jbi.12023. hdl:10261/123988. S2CID 86520333.

[Oldfield2014-4] Oldfield, R.G.; Mandrekar, K.; Nieves, M.X.; Hendrickson, D.A.; Chakrabarty, P.; Swanson, B.O.; Hofmann, H.A. (2015). "Parental care in the Cuatro Ciénegas cichlid, Herichthys minckleyi (Teleostei: Cichlidae)". Hydrobiologia. 748 (1): 233–257. doi:10.1007/s10750-014-2081-4. S2CID 17487888.

[5] López-Fernández H.; Winemiller K.O.; Honeycutt R.L. (2010). "Multilocus phylogeny and rapid radiations in Neotropical cichlid fishes (Perciformes: Cichlidae)". Molecular Phylogenetics and Evolution. 55 (3): 1070–1086. doi:10.1016/j.ympev.2010.02.020. PMID 20178851.

[6] Hans-Joachim P.; Morgenstern, R. & Schindler, I. (2014). "Cichlid fishes (Teleostei, Cichlidae) collected by Ferdinand Deppe in Mexico" (PDF). Vertebrate Zoology. 64 (1): 43–57. doi:10.3897/vz.64.e31462. S2CID 257187441. Archived from the original (PDF) on 2014-12-19. Retrieved 2014-12-19.

[Mcmahan_C._D.,_Matamoros_W._A.,_Piller_K._R.,_Chakrabarty_P._2015_211–234-7] Mcmahan C. D.; Matamoros W. A.; Piller K. R.; Chakrabarty P. (2015). "Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera". Zootaxa. 3999 (2): 211–234. doi:10.11646/zootaxa.3999.2.3. PMID 26623572. S2CID 34699955.

[CoF-8] Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Species in the genus Herichthys". Catalog of Fishes. California Academy of Sciences. Retrieved 10 November 2018.