Kota Formation

Kota Formation
Kota Formation
Stratigraphic range: Jurassic
Type	Geological formation
Sub-units	Lower & Upper members
Underlies	Unconformity with the Gangapur Formation
Overlies	Dharmaram Formation
Thickness	550–600 m (1,800–1,970 ft)
Lithology
Primary	Mudstone, sandstone
Other	Limestone
Location
Coordinates	18.9°N 80.0°E
Approximate paleocoordinates	31.6°S 31.9°E
Region	Telangana
Country	India
Extent	Pranhita-Godavari Basin
Type section
Named for	Kota Village
	Kota Formation (India)

The Kota Formation is a geological formation in India. The age of the Kota Formation is uncertain; it is commonly considered to date to the Early Jurassic, but some studies have suggested it may extend into the Middle Jurassic or even later. It conformably overlies the Lower Jurassic Upper Dharmaram Formation and is unconformably overlain by the Lower Cretaceous Gangapur Formation. It is split into a Lower Member and Upper Member. The Lower Member is approximately 100 m thick while the Upper Member is 490 m thick. Both subunits primarily consist of mudstone and sandstone, but near the base of the upper unit there is a 20-30 metre thick succession of limestone deposited in a freshwater setting.[1]

Stratigraphy

The lower boundary of the Kota Formation is made of pebbly sandstone, covering the topmost clay seen in the Dharmaram Formation.[2] The Kota Formation has been traditionally divided into 2 main members, the Lower and Upper members, yet more recent work have redivided it into 3.[3] The Lower member can be seen at locations such as Adamilli, Kamavarapukota and Sudikonda, being made of sandstones, with clay clasts, with greater or lower stratification.[4] The Middle Member is well developed along the Continental Gondwana basin, specially towards the northwestern part, and is made of medium to fine white sandstone with clay and concretionary limestone, suggesting the development of paleosols associated with alluvial floodplains.[3] The last member is mostly made of broad sandstone sheets with large clay casts associated with fluvial channels, and has an extension that can be easuly seen on several continuous kilometers.[4][5] The Uppermost section of the unit is mostly made of limestones and is overlain on an angular unconformity by the Gangapur Formation.[3]

Age

The age of the Kota Formation is controversial. There are no magmatic rocks or volcanic ash beds associated with the Kota Formation, which means that its age cannot be determined directly through radiometric dating.[6][7] The maximum age of the Kota Formation is constrained by the underlying Upper Dharmaram Formation, which is Early Jurassic, probably Hettangian or Sinemurian, in age.[8][7] Various researchers have attempted to date the Kota Formation using biostratigraphy. Krishnan (1968), Jain (1973), and Yadagiri and Prasad (1977) favored an Early Jurassic age based on the fish fauna. Govindan (1975) suggested a Middle Jurassic age based on ostracods. In 2006, Bandyopadhyay and Sengupta argued that the fish fauna suggested a Toarcian age for the Upper Kota Formation, possibly extending into the Aalenian, and in turn estimated the Lower Kota to be Sinemurian to Pliensbachian in age.[9] Guntupalli V. R. Prasad, along with various coauthors, has argued for a younger age. In 2001, Vijaya and Prasad proposed based on palynological evidence that the Kota Formation was deposited between the Callovian age of the Middle Jurassic and the Barremian age of the Early Cretaceous.[10] In 2002, Prasad and Manhas argued that the mammal genus Dyskritodon, known only from the Kota Formation and the Early Cretaceous of Morocco, provides evidence for a young age for the Kota Formation.[11] In 2020, Prasad and Parmar argued that the similarity of the dinosaur fauna of the Kota Formation to that of the Middle Jurassic of the United Kingdom supported a Middle Jurassic age for the Kota Formation.[12]

Paleoenvironment

The Kota Formation hosted lacustrine and wetland settings, this last one being of carbonate type, having modern analoges such as the Tablas de Daimiel in Spain

The Kota Formation represents mostly a Continental succession related to a continental rift basin, the Pranhita-Godavari Gondwana Basin of peninsular India.[13] The associated facies of sandstone and limestones are likely related to playa-type lake, with nearby fluvial currents, part of low gradient hanging wall alluvial fans, being deposited on it´s margin. There have been records of freshwater lue green algal stromatolites and oncolites, suggested to be deposited on low energy and low bathymetry lacustrine settings.[14] More recent works have proven the basin hosted in the Early Jurassic a freshwater carbonate wetland marked by the presence of limestones.[1] The environmental model proposed include a depositional cycle marked by several facies types, A for the sublittoral zones of shallow water bodies, followed by palustrine environments, including surfaces with abundance of influence of both plants and animals, specially rhizobrecciation indicating active colonization of the margins by plants, having a similar deposition to the modern Las Tablas de Daimiel wetlands.[1] Associated with the lacustrine facies have recovered microbial bioherms and lacustrine spring mounds, shallow ephemeral ponds with carbonated mud and Phyllopods, pedogenic calcrete under arid seasons and short-lived distributary channels.[13] The depositional setting may have been partially sheltered from the input of siliciclastic materials, except on flooding seasons. Microbial biomats likely developed on shallow waters, while rhizoliths increased it´s presence of abandoned channel fills and pedogenic facies indicate drought seasons.[13]

Fossil content

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Ostracoda

Ostracods
Genus	Species	Location	Stratigraphic position	Material	Notes
Clinocypris[15]	Clinocypris sp.	Daroghapalli	Upper Member	Six carapaces	A freshwater ostracodan of the Family Pontocyprididae.
Cypredea[15]	Cypredea sp.	Daroghapalli	Upper Member	Fourteen incomplete carapaces	A freshwater ostracodan of the Family Palaeocytheridae.
Darwinula[16][15]	Darwinula cf.sarytirmenensis	Aklapalli Daroghapalli Kanchelli Metpalli Potepalli	Lower Member Upper Member	More than 200 carapaces and valves	A freshwater ostracodan of the Family Darwinulidae. The most dominant genus locally and the main indicator of both fluvial and lacustrine settings
	Darwinula kingi	Aklapalli Daroghapalli Kanchelli Metpalli Potepalli	Lower Member Upper Member	Around 120 carapaces and valves	A freshwater ostracodan of the Family Darwinulidae.
	Darwinula spp.	Daroghapalli	Lower Member Upper Member	Nineteen Carapaces	A freshwater ostracodan of the Family Darwinulidae.
Eucandona[15]	Eucandona sp.	Daroghapalli	Upper Member	Eight incomplete carapaces	A freshwater ostracodan of the Family Candoninae.
Limnocythere[16]	Limnocythere spp.	Daroghapalli	Lower Member	Three complete carapaces	A freshwater ostracodan of the family Limnocytheridae.
Stenocypris[15]	?Stenocypris sp.	Daroghapalli	Upper Member	Single incomplete carapace	A freshwater ostracodan of the family Cyprididae.
Timiriasevia[16]	Timiriasevia digitalis	Daroghapalli	Lower Member	Twenty complete carapaces and thirty-six partly broken carapaces.	A freshwater ostracodan of the family Limnocytheridae.

Phyllopoda

Phyllopods
Genus	Species	Location	Stratigraphic position	Material	Notes
Estheriina[17][18]	Estheriina alibadadensis	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Estheriininae. The most abundant Estheriid in the region and the key element of the Estheriina biozone
	Estheriina indijurassica	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Estheriininae.
	Estheriina bullata	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Estheriininae.
	Estheriina pranhitaensis	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Estheriininae.
Lioestheria[17][18]	Lioestheria kotaensis	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Lioestheriidae. The second key element of the Estheriina biozone
	Lioestheria crustabundis[17]	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Lioestheriidae.
	Lioestheria ssp.[17]	Kota limestone ridge	Lower Member	Valves	A freshwater clam shrimp of the family Lioestheriidae.
Paleolimnadia[17]	Paleolimnadia spp.	Kota limestone ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Estheriininae.
Pseudeasmussiata[17]	Pseudeasmussiata andhrapradeshia	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	Valves	A freshwater clam shrimp of the family Lioestheriidae.

Insecta

Insects
Genus	Species	Location	Stratigraphic position	Material	Notes
Archimesoblatta[19]	A. shiva	Tasch outcrop K-2 bed 8	Upper Member	Right forewing tegmen	A mesoblattinid cockroach
Coleopteron[20]	C. sp.	Kota limestone ridge	Lower Member	Isolated wings	An Indeterminate Coleopteran.
Kotaphialtites[21]	K. frankmortoni	Sirpur Taluka, Tasch's K1 outcrop bed 2(A)	Upper Member	MCZ 11909, Isolated wing	An ephialtitid hymenopteran.
Protogryllus[22]	P. lakshmi	Kota Formation outcrop K-2	Upper Member	MCZ 3046, Isolated wing	A protogryllid cricket
Taschigatra[23]	T. bharataja	Sirpur Taluka, Tasch's K1 outcrop bed 3(A)	Upper Member	No.2013(3013), part and counterpart of well preserved wing	A rhagionid dipteran.
Taschigatra[23]	T. tulyabhijana	Sirpur Taluka, Tasch's K1 outcrop bed 2(A)	Upper Member	No. 5034, well preserved wing	A rhagionid dipteran.
Xyelula[21]	X. alexandri	Kota Formation outcrop K-2	Upper Member	MCZ 11831, well preserved wing	A sepulcid hymenopteran.

Fish

Fish
Taxon	Species	Location	Stratigraphic position	Material	Notes
Indocoelacanthus[24]	Indocoelacanthus robustus	Boraigudem limestone ridge Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	ISI P. 39. Impression of the caudal fin ISI P. 40. Nearly complete head and trunk squamation ISI P. 41-45. Fragments of fin-rays ISI P. 46. Neural arches, isolated scales and girdle elements	A robust freshwater coelacanth of the family Latimeriidae. Represents the largest member of the local freshwater fauna, measuring up to 70 cm.[24]
Lepidotes[25][26][27]	Lepidotes deccanensis	Kota limestone ridge Paikasigudem village	Lower Member Upper Member	Complete Specimen Isolated Remains	A freshwater neopterygian of the family Lepisosteiformes.
Lepidotes[25][26][27]	Lepidotes spp.	Kota limestone ridge Paikasigudem village	Lower Member Upper Member	Isolated remains	A freshwater neopterygian of the family Lepisosteiformes.
Lonchidion[28][25]	Lonchidion indicus	Paikasigudem village Yamanpalli bonebed	Upper Member	GSI.TI.2, tooth Isolated Teeth	A freshwater elasmobranch of the family Lonchidiidae.
Paradapedium[25][29]	Paradapedium egertoni	Boraigudem limestone ridge Kota limestone ridge Near Ankisha	Lower Member Upper Member	BMNHP12146, Partial specimen BMNHP12147, Almost complete specimen BMNHP12148, Partial specimen ISIP.32 Nearly complete specimen ISIP.33 Nearly complete specimen ISIP.34 Nearly complete specimen ISIP.35 specimen lacking tail and head	A freshwater neopterygian of the family Dapediidae.
Pholidophorus[30]	Pholidophorus kingi	Boraigudem limestone ridge Kota limestone ridge Paikasigudem village	Lower Member Upper Member	Several complete and incomplete specimens	A freshwater neopterygian of the family Pholidophoridae.
Pholidophorus[30]	Pholidophorus indicus	Boraigudem limestone ridge Kota limestone ridge Paikasigudem village	Lower Member Upper Member	Several complete and incomplete specimens	A freshwater neopterygian of the family Pholidophoridae.
Polyacrodus[25][31]	Polyacrodus? sp.	Paikasigudem village	Upper Member	Tooth	A freshwater elasmobranch of the family Polyacrodontidae.
Pycnodontidae[28]	Indeterminate	Paikasigudem village	Upper Member	About 60 specimens of Pharyngeal Teeth, including GSITL3-6	A freshwater neopterygian of the family Pycnodontidae, originally classified as Perciformes, yet suggested to be very similar to the Cretaceous pycnodont Stephanodus.
Semionotiformes[25]	Indeterminate	Paikasigudem village	Upper Member	Teeth	A freshwater neopterygian of the family Semionotiformes.
Tetragonolepis[25][29]	Tetragonolepis oldhami	Kota limestone ridge Lingal-Metpalli ridge	Lower Member Upper Member	GSI2145, an impression including most of the body ISP36, fragment of trunk and skull ISP37, nearly complete specimen	A freshwater neopterygian of the family Dapediidae.

Amphibia

Amphibians
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Anura[28]	Indeterminate	Paikasigudem village	Upper Member	GSI.TL.19, Mandible GSI.TL.20, right illium	Indeterminate frog remains, originally referred to Pelobatidae due to be compared with younger Creteaceous Indian frog material
Caudata[28]	Indeterminate	Paikasigudem village	Upper Member	GSI.TI.10, mandible with one teeth GSI.TI.16, fragment of mandible GSI.TI.17, madible with 4 teeth	Indeterminate caudatan remains, originally referred to Sirenidae due to be compared with younger Creteaceous sirenid material

Mammaliaforms

Mammaliaforms
Taxon	Species	Location	Stratigraphic position	Material	Notes
Australosphenida[32]	Indeterminate	Paikasigudem village	Upper Member	Isolated lower molar	A mammal of the group Australosphenida, resembling the south american genus Asfaltomylos
Dyskritodon?[11]	Dyskritodon? indicus	Paikasigudem village	Upper Member	VPL/JU/KM/13, lower left molar	A dubious mammal of the group Eutriconodonta. This Genus is known from the Early Cretaceous of Morocco, what has been used to suggest a minimum Berrasian age for the Upper Kota Formation
Gondtherium[31]	Gondtherium dattai	Paikasigudem village	Upper Member	VPL/JU/KM 12 right lower molar	A mammal of the family Docodontidae
Indotherium[33]	Indotherium pranhitai	5 km west of Yamanapalli	Lower Member	GSI20795, right upper molar	A mammaliform of the family Morganucodontidae. Includes the informally named "Indozostrodon simpsoni".[34]
Indobaatar[35]	Indobaatar zofiae	Paikasigudem village	Upper Member	VPL/JU/KM/20, a left upper premolar	A mammal described as an eobaatarid multituberculate, but this interpretation has been challenged.[36]
Kotatherium[37]	Kotatherium haldanei	5 km west of Yamanapalli	Lower Member	GSI19634, right upper molar	A mammaliform of the family Kuehneotheriidae
Nakunodon[38]	Nakunodon paikasiensis	Paikasigudem village	Upper Member	GSI.SR/PAL/12, right upper molar	A mammal of the family Amphidontidae
Paikasigudodon[11][39]	Paikasigudodon yadagirii	Paikasigudem village	Upper Member	VPL/JU/KM/10, right upper molar	A mammaliaform of the family Morganucodontidae, originally known as "Kotatherium yadagirii"
Trishulotherium[33]	Trishulotherium kotaensis	Paikasigudem village	Upper Member	GSISR/PAL/10, left lower molar	A mammal of the order Symmetrodonta

Lepidosauromorpha

Lepidosauromorphs
Taxon	Species	Location	Stratigraphic position	Material	Notes
Bharatagama[40]	Bharatagama rebbanensis	Paikasigudem village	Upper Member	VPL/JU/KR 66, dentary VPL/JU/KR 88, anterior region of a left maxilla VPL/JU/KR 91, left maxilla VPL/JU/KR 90-92, posterior ends of right maxillae VPL/JU/KR 67-80-103, anterior symphysial region JU/KR 79, 98, 100, dentaries mid-region VPL/JU/KR 69, 81-84, 87, dentary with hatchling dentition VPL/JU/KR 68, 85-86, dentary VPL/JU/KR 70, 71, 76,78, 93, 94, 97, 104, 105, dentary	A lepidosauromorph originally described as an Iguanian lizard. May actually be a sphenodontian rather than a lizard.[41]
Godavarisaurus[42]	Godavarisaurus lateefi	Gorlapalli village Paikasigudem village	Upper Member	VP4Ju/KR47, a partial right dentary VP4Ju/KR44, 51, 52, right maxilla VP4Ju/KR40, 41, 43, 49, 50, left maxilla VP4Ju/KR37, 48, 65, right dentary VP4Ju/KR39, fragment of left dentary VP4Ju/KR46, left palatine	A small sphenodontian, with a skull estimated to measure less than 20 mm
Paikasisaurus[28][40]	Paikasisaurus indicus	Paikasigudem village	Upper Member	GSI.TI.14, left dentary with teeth GSI.TI.15, left dentary with one tooth	An indeterminate and dubious lepidosauromorph, originally suggested to be a varanoid lizard
Rebbanasaurus[42]	Rebbanasaurus jaini	Paikasigudem village	Upper Member	About 50 specimens, including premaxillae, maxillae, dentaries, and a palatine	A small sphenodont
Squamata?[28][40]	Indeterminate	Paikasigudem village	Upper Member	VPL/JU/KR 62, fragmentary right maxilla VPL/JU/KR 61, partial maxilla	Distinct from Bharatagama rebbanensis; may include material formerly assigned to the dubious Kota squamate Paikasisaurus indicus.[lower-alpha 1]
Sphenodontidae[28][42]	Indeterminate	Paikasigudem village	Upper Member	GSI.TI.8-12, 17, maxilla	Indeterminate sphenodontidae remains

Testudinata

Turtles
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Indochelys[43][44]	Indochelys spatulata	3 km NNE of Kistapur village, Kota Near Kota village, north of Sironcha	Upper Member	GSI 20380, a partial shell ISI R176, a partial shell ISI R177, the anterior half of a shell	Mesochelydian stem-turtle, suggested to be related with Condorchelys
Testudines[45]	Indeterminate	1 km south of Bodepalli	Lower Member	Carapace fragments	Indeterminate turtle remains

Crocodylomorpha

Atoposauridae crocodiles are known from the unit, yet is not clear from what locality.[7]

Crocodylomorpha
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Crocodylomorpha[46]	Indeterminate	1 km south of Bodepalli	Lower Member	Maxillae, dentaries, teeth	Indeterminate crocodylomorph remains, previously mixed with thyreophoran material and part of the chimaeric "Andhrasaurus"
Teleosauridae?[47]	Indeterminate	Kota limestone ridge	Lower Member	Dermal scutes, with a femur and some fragments of other bones	Indeterminate material referred to crocodylomorphs similar to Teleosaurus

Pterosauria

Pterosaurs
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Campylognathoides[48]	Campylognathoides indicus	Kota limestone ridge	Lower Member	ISI R38, holotype, fragment of skull and upper jaw	The holotype of Campylognathoides indicus, a pair of premaxillae, may represent a fish rather than a pterosaur.[49]
Pterosauria[48][50]	Indeterminate	Boraigudem Chitur	Lower Member	GSI17868, radius and ulna, wing metacarpal, first wing-phalanx and a clawed toe ISIR49. fragment displaying dissociated bones from post-cranial region	Indeterminate pterosaur remains

Dinosaurs

Dinosaurs
Genus	Species	Location	Stratigraphic position	Material	Notes
"Andhrasaurus"[45][51]	"Andhrasaurus indicus"	1 km south of Bodepalli	Lower Member	Sacral vertebra, vertebral centra, dorsal vertebrae, caudal vertebrae, parts of scapula and ilium, osteoderms	A chimaera of thyreophoran postcranial material and Crocodrylomorph skull pieces. The armor was later suggested to be Ankylosauria indet.[46] And other latter works pointed out it likely belongs to an indeterminate basal thyreophoran.[52]
Barapasaurus[53]	Barapasaurus tagorei	North of Krishnapur Pochampalli village	Lower Member	Roughly 300 bones, representing the disarticulated remains of at least six individuals	A sauropod dinosaur, either a Eusauropoda or more likely a Gravisauria. Represents the best-known Early Jurassic sauropod
"Dandakosaurus"[54]	"Dandakosaurus indicus"	Yamanpalli bonebed	Lower Member	Lateral tooth, proximal ischium Dorsal vertebra, proximal caudal vertebra (Likely of Kotasaurus)	A chimaera of large theropod bones, including and ischium and tooth, probably belonging to a carnosaur, and sauropod bones (two Kotasaurus vertebrae)
Dromaeosauridae?[12]	Indeterminate	Paikasigudem village	Upper Member	Isolated Teeth	Five distinct morphotypes have been identified, mostly resemble coelurosaurs or dromaeosauroids.[12]
Hypsilophodontidae[55]	Indeterminate	Gorlapalli Village	Upper Member	Single Tooth	Identified originally as a member of Hypsilophodontidae, probably represents a tooth of a basal neornithischian.[12]
Kotasaurus[56]	Kotasaurus yamanpalliensis	Yamanpalli bonebed	Lower Member	Disarticulated remains of at least 12 individuals[56]	A basal sauropod
Ornithischia[46][12]	Indeterminate	1 km south of Bodepalli Paikasigudem village	Lower Member Upper Member	End of left humerus Isolated Teeth	Indeterminate ornithischian material. Among the teeth, at least five distinct morphotypes have been identified.
Richardoestesia?[12]	Richardoestesia? spp.	Paikasigudem village	Upper Member	Isolated Teeth	Teeth similar to those of the problematic taxon Richardoestesia, of supposed coelurosaur affinities
Scelidosauridae[12][57]	Indeterminate	Paikasigudem village	Upper Member	Scute and associated fragmentary limb bones	Indeterminate scelidosaurid material

Megaflora

Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Agathoxylon[58]	Agathoxylon kotaense	Near Kota village	Lower Member	Fossil wood	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Agathoxylon chandrapurensis[59]	Near Kota village	Lower Member	Fossil wood	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Agathoxylon santacruzense[60]	Near Kota village	Lower Member	Fossil wood	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Agathoxylon pranhitaensis[58][61]	Near Kota village	Lower Member	Fossil wood	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Agathoxylon santalense[58]	Near Kota village	Lower Member	Fossil wood	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Agathoxylon spp.[58]	Bodepalli Near Kota village Paikasigudem village	Lower Member Upper Member	Fossil wood	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Araucarites[61][50]	Araucarites minutus	Dronadula	Lower Member	Branched shoots	Affinities with Araucariaceae inside Pinales.
Araucarites[61][50]	Araucarites sp.	Near Kota village	Lower Member	Branched shoots	Affinities with Araucariaceae inside Pinales.
Brachyphyllum[61][59]	Brachyphyllum expansum	Near Kota village	Lower Member	Branched shoots	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Elatocladus[61][50]	Elatocladus conferta	Near Kota village	Lower Member	Branched shoots	Affinities with Cupressaceae inside Pinales.
	Elatocladus tenerrimus[59]	Near Kota village	Lower Member	Branched shoots	Affinities with Cupressaceae inside Pinales.
	Elatocladus jabalpurensis[59]	Near Kota village	Lower Member	Branched shoots	Affinities with Cupressaceae inside Pinales.
	Elatocladus plana[59]	Near Kota village	Lower Member	Branched shoots	Affinities with Cupressaceae inside Pinales.
	Elatocladus sp.[59]	Near Kota village	Lower Member	Branched shoots	Affinities with Cupressaceae inside Pinales.
Cladophlebis[50]	Cladophlebis denticulata	Boraigudem Chitur Near Kota village	Lower Member	Isolated fronds	Affinities with Osmundaceae in the Osmundales.	Cladophlebis denticulata illustration
	Cladophlebis indica[61][50]	Boraigudem Chitur Near Kota village	Lower Member	Isolated fronds	Affinities with Osmundaceae in the Osmundales
	Cladophlebis reversa[61][50]	Boraigudem Chitur Near Kota village	Lower Member	Isolated fronds	Affinities with Osmundaceae in the Osmundales
	Cladophlebis spp.[61][50]	Boraigudem Chitur Near Kota village	Lower Member	Isolated fronds	Affinities with Osmundaceae in the Osmundales
Circoporoxylon[62]	Circoporoxylon kotaense	Near Kota village	Lower Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
Coniopteris[61][50]	Coniopteris hymenophylloides	Boraigudem Chitur Near Kota village	Lower Member	Isolated fronds	Affinities with Polypodiales in the Polypodiidae. Common cosmopolitan Mesozoic fern genus. Recent research has reinterpreted it a stem group of the Polypodiales (closely related to the extant genera Dennstaedtia, Lindsaea, and Odontosoria)	Coniopteris specimen
Coniopteris[61][50]	Coniopteris sp.[61]	Near Kota village	Lower Member	Isolated fronds	Affinities with Polypodiales in the Polypodiidae.	Coniopteris specimen
Cupressinoxylon[58][61]	Cupressinoxylon kotaense	Near Kota village	Lower Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
Dictyozamites[59]	Dictyozamites falcatus	Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
Dictyozamites[59]	Dictyozamites kotaense[59]	Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
Equisetum[61]	Equisetum rajmahalensis	Chitur village Near Kota village	Lower Member Upper Member	Isolated Stems	Affinities with Equisetaceae inside Equisetales.	Extant Equisetum arvense specimen
Ginkgoites[61]	Ginkgoites lobata	Near Kota village	Lower Member	Leafs	Affinities with Ginkgoaceae inside Ginkgoopsida.	Ginkgoites reconstruction
Ginkgoxylon[63]	Ginkgoxylon dixii	Near Kota village	Lower Member	Fossil wood	Affinities with Ginkgoaceae inside Ginkgoopsida.
Hausmannia[61]	Hausmannia cf. buchii	Near Kota village	Lower Member	Isolated pinnae	Affinities with Dipteridaceae in the Polypodiales.	Hausmannia specimen
Otozamites[59]	Otozamites vemavarmensis	Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.	Otozamites specimen
Pagiophyllum[61][50]	Pagiophyllum peregrinum	Near Kota village	Lower Member	Branched shoots	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Pagiophyllum cf.peregrinum	Near Kota village	Lower Member	Branched shoots	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
	Pagiophyllum spp.	Near Kota village	Lower Member	Branched shoots	Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.
Pachypteris[59]	Pachypteris indica	Near Kota village	Lower Member	Isolated pinnae	Affinities with Umkomasiaceae in the Pteridospermatophyta.
Planoxylon[59]	Planoxylon mahabalei	Near Kota village	Lower Member	Fossil wood	Affinities with Protopinaceae inside Pinales.
Podocarpoxylon[61]	Podocarpoxylon chandrapurensis	Near Kota village	Lower Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
	Podocarpoxylon chiturensis[59]	Chitur village	Upper Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
	Podocarpoxylon krauselii[61]	Near Kota village	Lower Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
	Podocarpoxylon rajmahalense[61]	Near Kota village	Lower Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
	Podocarpoxyion sewardii[59]	Near Kota village	Lower Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
	Podocarpoxylon sp.[61]	Near Kota village	Lower Member	Fossil wood	Affinities with Podocarpaceae inside Pinales.
Podozamites[61]	Podozamites sp.	Near Kota village	Lower Member	Leaflets	Broad conifer leaves
Pseudoctenis[59]	Pseudoctenis cf. frngilis	Near Kota village	Lower Member	Leaflets	Affinities with Cycadales in the Cycadopsida.
Ptilophyllum[61]	Ptilophyllum fissum	Boraigudem Chitur Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
	Ptilophyllum acutifolium	Boraigudem Chitur Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
	Ptilophyllum cutchense[59]	Chitur village Near Kota village	Lower Member Upper Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
	Ptilophyllum cf.sahnii[59]	Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
	Ptilophyllum cf.institacallum[59]	Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
	Ptilophyllum sp.	Near Kota village	Lower Member	Leaflets	Affinities with Williamsoniaceae in the Bennettitales.
Protaxodioxylon[64]	Protaxodioxylon sahnii	Chitur village	Upper Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
Protaxodioxylon[64]	Prototaxoxylon liassicum[65]	Near Kota village	Lower Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
Sphenopteris[61][59]	Sphenopteris kotaensis	Near Kota village	Lower Member	Isolated Fronds	Affinities with Dicksoniaceae in the Cyatheales.
Taxaceoxylon[61]	Taxaceoxylon sahnii	Near Kota village	Lower Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
	Taxaceoxylon biradarii[59]	Chitur village	Lower Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
	Taxaceoxylon antiquum[59]	Near Kota village	Lower Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
	Taxaceoxylon sp.	Near Kota village	Upper Member	Fossil wood	Affinities with Cupressaceae inside Pinales.
Torreyites[59]	Torreyites constricte	Chitur village	Upper Member	Branched shoots	Affinities with Taxaceae in the Pinales.	Extant Torreya specimen

Notes

It cannot be determined which of the two Kota squamate forms, if either, the holotype of Paikasisaurus indicus pertains to.[40]

References

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[43] It cannot be determined which of the two Kota squamate forms, if either, the holotype of Paikasisaurus indicus pertains to.[40]

[Sedi-1] Goswami, Suparna; Gierlowski-Kordesch, Elizabeth; Ghosh, Parthasarathi (January 2018). "Sedimentology of the Early Jurassic limestone beds of the Kota Formation: record of carbonate wetlands in a continental rift basin of India". Journal of Paleolimnology. 59 (1): 21–38. Bibcode:2018JPall..59...21G. doi:10.1007/s10933-016-9918-y. ISSN 0921-2728. S2CID 133167210.

[2] Kutty, T. S.; Jain, S. L.; Chowdhury, T. R. (1987). "Gondwana sequence of the northern Pranhita-Godavari Valley: its stratigraphy and vertebrate faunas" (PDF). Palaeobotanist. 36: 263–282. Retrieved 16 June 2023.

[MemberAA-3] Lakshminarayana, G. (1994). "Stratigraphy and structural framework of the Gondwana sediments in the Pranhita-Godavari Valley, Andhra Pradesh". Gondwana Nine. 1 (2): 311–330.

[MemberAB-4] Lakshminarayana, G. (2002). "Evolution in basin fill style during the Mesozoic Gondwana continental break-up in the Godavari Triple junction, SE India". Gondwana Research. 5 (1): 227–244. Bibcode:2002GondR...5..227L. doi:10.1016/S1342-937X(05)70906-0. Retrieved 16 June 2023.

[5] Sengupta, S. (2003). "Gondwana sedimentation in the Pranhita–Godavari Valley: a review". Journal of Asian Earth Sciences. 21 (6): 633–642. Bibcode:2003JAESc..21..633S. doi:10.1016/S1367-9120(02)00052-4. Retrieved 16 June 2023.

[Chinnappa2019-6] Chinnappa, Chopparapu; Rajanikanth, Annamraju; Pauline Sabina, Kavali (2019). "Palaeofloras from the Kota Formation, India: palaeodiversity and ecological implications". Volumina Jurassica. 17: 1–16. doi:10.7306/vj.17.1 (inactive 1 August 2023).{{cite journal}}: CS1 maint: DOI inactive as of August 2023 (link)

[Chatterjee2020-7] Chatterjee, Sankar (2020). "The Age of Dinosaurs in the Land of Gonds". In Prasad, Guntupalli V.R.; Patnaik, Rajeev (eds.). Biological Consequences of Plate Tectonics. Vertebrate Paleobiology and Paleoanthropology. Cham: Springer International Publishing. pp. 181–226. doi:10.1007/978-3-030-49753-8_8. ISBN 978-3-030-49752-1. S2CID 229651571.

[Kutty2007-8] Kutty, T.S.; Chatterjee, Sankar; Galton, Peter M.; Upchurch, Paul (2007). "Basal sauropodomorphs (Dinosauria: Saurischia) from the Lower Jurassic of India: their anatomy and relationships". Journal of Paleontology. 81 (6): 1552–1574. Bibcode:2007JPal...81.1218K. doi:10.1666/04-074.1. S2CID 130508134.

[Bandyopadhyay2006-9] Bandyopadhyay, Saswati; Sengupta, Dhurjati Prasad (2006). "Vertebrate faunal turnover during the Triassic-Jurassic transition: an Indian scenario". New Mexico Museum of Natural History and Science Bulletin. 37: 77–85.

[Vijaya2001-10] Vijaya; Prasad, G. V. R. (2001). "Age of the Kota Formation, Pranhita-Godavari Valley, India: a palynological approach". Journal of the Palaeontological Society of India. 46: 77–93.

[Prasad2002-11] Prasad, Guntupalli V R; Manhas, Brijesh K (2002). "Triconodont mammals from the Jurassic Kota Formation of India". Geodiversitas. 24 (2): 445–464.

[Prasad2020-12] Prasad, Guntupalli V. R.; Parmar, Varun (2020). "First Ornithischian and Theropod Dinosaur Teeth from the Middle Jurassic Kota Formation of India: Paleobiogeographic Relationships". In Prasad, Guntupalli V.R.; Patnaik, Rajeev (eds.). Biological Consequences of Plate Tectonics. Cham: Springer International Publishing. pp. 1–30. ISBN 978-3-030-49752-1.

[Micro-13] Goswami, S.; Ghosh, P. (2021). "Freshwater Microbialites in Early Jurassic Fluvial Strata of the Pranhita-Godavari Gondwana Basin, India". _Progress, Challenges and Opportunities. Syntheses in Limnogeology: 549–578. doi:10.1007/978-3-030-66576-0_18. ISBN 978-3-030-66575-3. S2CID 237995280. Retrieved 16 June 2023.

[14] Rudra, D. K.; Maulik, P. K. (1987). "Stromatolites from Jurassic freshwater limestone, India". Mesozoic Research. 1 (3): 135–146.

[Ostrac2-15] Misra, R.S.; Satsangi, P.P. (1979). "Ostracods from Kota Formation". Proceedings of the Colloquium on Micropalaeontology and Stratigraphy. Geological Survey of India, Miscellaneous Publication. 45 (5): 73–80.

[Ostrac1-16] Govindan, A. (1975). "Jurassic fresh water ostracods from the Kota limestones of India". Palaeontology. 18 (4): 207–216. Retrieved 15 June 2023.

[Esthe1-17] Tasch, P. (1987). Fossil Conchostraca of the Southern Hemisphere and continental drift: Paleontology, biostratigraphy, and dispersal (165 ed.). Geological Society of America.

[Estheria-18] Ghosh, S. C.; Datta, A.; Nandi, A.; Mukhopadhyay, S. (1987). "Estheriid zonation in the Gondwana" (PDF). Paleobotanist: 99–123. Retrieved 16 June 2023.

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