Nemastomatinae

Nemastomatinae is a subfamily of harvestmen with 123 described species in 18 genera.

Nemastomatinae
Nemastoma lugubre
Scientific classification
Kingdom:
Animalia
Phylum:
Arthropoda
Class:
Arachnida
Order:
Opiliones
Suborder:
Dyspnoi
Superfamily:
Ischyropsalidoidea
Family:
Nemastomatidae
Subfamily:
Nemastomatinae

Simon, 1872
Genera
Diversity
18 genera, 123 species

Species

The following species are included in Nemastomatinae:[1]

  • Acromitostoma Roewer, 1951
  • Acromitostoma hispanum (Roewer, 1917)
  • Acromitostoma rhinoceros (Roewer, 1917)
  • Carinostoma Kratochvíl, 1958
  • Carinostoma carinatum (Roewer, 1914)
  • Carinostoma elegans (Sørensen, 1894)
  • Carinostoma ornatum (Hadži, 1940)
  • Caucnemastoma Martens, 2006
  • Caucnemastoma golovatchi Martens, 2006
  • Caucnemastoma martensi Snegovaya, 2011
  • Centetostoma Kratochvíl, 1958
  • Centetostoma centetes (Simon, 1881)
  • Centetostoma juberthiei Martens, 2011
  • Centetostoma scabriculum (Simon, 1879)
  • Centetostoma ventalloi (Mello-Leitao, 1936)
  • Giljarovia Kratochvíl, 1958
  • Giljarovia crimeana Chemeris & Kovblyuk, 2012[2]
  • Giljarovia kratochvili Snegovaya, 2011
  • Giljarovia redikorzevi Charitonov, 1946
  • Giljarovia rossica Kratochvíl, 1958
  • Giljarovia stridula (Kratochvíl, 1958)
  • Giljarovia tenebricosa (Redikortsev, 1936)
  • Giljarovia thoracocornuta Martens, 2006
  • Giljarovia triangula Martens, 2006
  • Giljarovia trianguloides Martens, 2006
  • Giljarovia turica Gruber, 1976
  • Giljarovia vestita Martens, 2006
  • Hadzinia Šilhavý, 1966
  • Hadzinia ferrani Novak & Kozel, 2014[3]
  • Hadzinia karamani (Hadži, 1940)
  • Histricostoma Kratochvíl, 1958
  • Histrocostoma anatolicum (Roewer, 1962)
  • Histricostoma argenteolunulatum (Canestrini, 1875)
  • Histricostoma caucasicum (Redikortsev, 1936)
  • Histricostoma creticum (Roewer, 1927)
  • Histricostoma dentipalpe (Ausserer, 1867)
  • Histricostoma drenskii Kratochvíl, 1958
  • Histricostoma gruberi Snegovaya & Marusik, 2012
  • Histricostoma mitovi Snegovaya & Marusik, 2012
  • Mediostoma Kratochvíl, 1958
  • Mediostoma armatum Martens, 2006
  • Mediostoma ceratocephalum Gruber, 1976
  • Mediostoma cypricum (Roewer, 1951)
  • Mediostoma globuliferum (L. Koch, 1867)
  • Mediostoma haasi (Roewer, 1953)
  • Mediostoma humerale (C.L. Koch, 1839)
  • Mediostoma izmirica Snegovaya, Yağmur & Kurt, 2016[4]
  • Mediostoma nigrum Martens, 2006
  • Mediostoma pamiricum Staręga, 1986
  • Mediostoma stussineri (Simon, 1885)
  • Mediostoma variabile Martens, 2006
  • Mediostoma vityae (Roewer, 1927)
  • Mitostoma Roewer, 1951
  • Mitostoma alpinum (Hadži, 1931)
  • Mitostoma anophthalmum (Fage, 1946)
  • Mitostoma atticum (Roewer, 1927)
  • Mitostoma cancellatum (Roewer, 1917)
  • Mitostoma carneluttii Hadži, 1973
  • Mitostoma chrysomelas (Hermann, 1804)
  • Mitostoma daccordii Tedeschi & Sciaky, 1997
  • Mitostoma fabianae Tedescie & Sciaky, 1997
  • Mitostoma gracile (Redikortsev, 1936)
  • Mitostoma macedonicum Hadži, 1973
  • Mitostoma olgae (Šilhavý, 1939)
  • Mitostoma olgae olgae (Šilhavý, 1939)
  • Mitostoma olgae decorum (Šilhavý, 1939)
  • Mitostoma olgae kratochvili (Šilhavý, 1939)
  • Mitostoma olgae zorae Hadži, 1973
  • Mitostoma orobicum (Caporiacco, 1949)
  • Mitostoma patrizii Roewer, 1953
  • Mitostoma pyrenaeum (Simon, 1879)
  • Mitostoma sabbadinii Tedeschi & Sciaky
  • Mitostoma valdemonense Marcellino, 1974
  • Mitostoma zmajevicae Hadži, 1973
  • Nemaspela Šilhavý, 1966
  • Nemaspela abchasica (Ljovuschkin & Starobogatov, 1963)
  • Nemaspela birsteini Ljovuschkin, 1972
  • Nemaspela caeca (Grese, 1911)
  • Nemaspela femorecurvata Martens, 2006
  • Nemaspela gagrica Chemeris, 2013[5]
  • Nemaspela kovali Chemeris, 2009
  • Nemaspela ladae Karaman, 2013[6]
  • Nemaspela sokolovi (Ljovuschkin & Starobogatov, 1963)
  • Nemaspela taurica (Lebedinski, 1914)
  • Nemastoma (C.L. Koch, 1836)
  • Nemastoma bidentatum Roewer, 1914
  • Nemastoma bidentatum bidentatum Roewer, 1914
  • Nemastoma bidentatum pluridentatum Hadži, 1973
  • Nemastoma bidentatum sparsum Gruber & Martens, 1968
  • Nemastoma bidentatum relictum Gruber & Martens, 1968
  • Nemastoma bimaculatum (Fabricius, 1775)
  • Nemastoma dentigerum Canestrini, 1873
  • Nemastoma lugubre (Müller, 1776)
  • Nemastoma schuelleri Gruber & Martens
  • Nemastoma transsylvanicum Gruber & Martens, 1968
  • Nemastoma triste (C.L. Koch, 1835)
  • Nemastomella Mello-Leitão, 1936
  • Nemastomella armatissima (Roewer, 1962)
  • Nemastomella bacillifera (Simon, 1879)
  • Nemastomella bacillifera bacillifera (Simon, 1879)
  • Nemastomella bacillifera carbonaria (Simon, 1907)
  • Nemastomella cristinae (Rambla, 1969)
  • Nemastomella dentipatellae (Dresco, 1967)
  • Nemastomella dipentata (Rambla, 1959)
  • Nemastomella dubia (Mello-Leitão, 1936)
  • Nemastomella gevia Prieto, 2004
  • Nemastomella hankiewiczii (Kulczyński, 1909)
  • Nemastomella iberica (Rambla in Dresco, 1967)
  • Nemastomella maarebensis (Simon, 1913)
  • Nemastomella manicata (Simon, 1913)
  • Nemastomella monchiquensis (Kraus, 1961)
  • Nemastomella sexmucronata (Simon, 1911)
  • Nemastomella spinosissima (Kraus, 1961)
  • Paranemastoma Redikorzev, 1936
  • Paranemastoma ancae Avram, 1973
  • Paranemastoma armatum (Kulczyński, 1909)
  • Paranemastoma aurigerum (Roewer, 1951)
  • Paranemastoma aurigerum aurigerum (Roewer, 1951)
  • Paranemastoma aurigerum ryla (Roewer, 1951)
  • Paranemastoma aurigerum joannae Staręga, 1976
  • Paranemastoma aurosum (L. Koch, 1869)
  • Paranemastoma beroni Mitov, 2011
  • Paranemastoma bicuspidatum (C.L. Koch, 1835)
  • Paranemastoma bureschi (Roewer, 1926)
  • Paranemastoma corcyraeum (Roewer, 1917)
  • Paranemastoma filipes (Roewer, 1917)
  • Paranemastoma iranicum Martens, 2006
  • Paranemastoma kalischevskyi (Roewer, 1951)
  • Paranemastoma kochii (Nowicki, 1870)
  • Paranemastoma longipes (Schenkel, 1947)
  • Paranemastoma quadripunctatum (Perty, 1833)
  • Paranemastoma radewi (Roewer, 1926)
  • Paranemastoma sillii (Herman, 1871)
  • Paranemastoma sillii sillii (Herman, 1871)
  • Paranemastoma sillii monticola Babalean, 2011
  • Paranemastoma simplex (Giltay, 1932)
  • Paranemastoma superbum Redikorzev, 1936
  • Paranemastoma thessalum (Simon, 1885)
  • Paranemastoma titaniacum (Roewer, 1914)
  • Paranemastoma werneri (Kulczyński, 1903)
  • Pyza Staręga, 1976
  • Pyza anatolica (Roewer, 1959)
  • Pyza bosnica (Roewer, 1917)
  • Pyza navarrense (Roewer, 1951)
  • Pyza taurica Gruber, 1979
  • Saccarella Schönhofer & Martens, 2012
  • Saccarella schilleri Schönhofer & Martens, 2012
  • Sinostoma Martens, 2016[7]
  • Sinostoma yunnanicum Martens, 2016
  • Starengovia Snegovaya, 2010
  • Starengovia kirgizica Snegovaya, 2010
  • Starengovia ivanloebli Martens, 2017[8]
  • Starengovia quadrituberculata Zhang & Martens, 2018[9]
  • Vestiferum Martens, 2006
  • Vestiferum alatum Martens, 2006
  • Vestiferum funebre (Redikorzev, 1936)

Fossil species

There are currently 5 described fossil harvestmen that have been assigned to Nemastomatinae, 4 of which are assigned to modern genera:[10]

  • Histricostoma Kratochvíl, 1958
  • † Histricostoma tuberculatum (C.L. Koch & Berendt, 1854) - Paleogene Baltic & Bitterfield amber
  • Mitostoma Roewer, 1951
  • † Mitostoma denticulatum (C.L. Koch & Berendt, 1854) - Paleogene Baltic amber
  • † Mitostoma gruberi Dunlop & Mitov, 2009 - Paleogene Baltic & Bitterfield amber
  • Nemastoma (C.L. Koch, 1836)
  • † Nemastoma incertum C.L. Koch & Berendt, 1854 - Paleogene Baltic amber
  • † Paragilarovia Elsaka, Mitov, & Dunlop, 2019[11]
  • † Paragilarovia hochae Elsaka, Mitov, & Dunlop, 2019 - Paleogene Baltic amber

References
  1. "A taxonomic catalogue of the Dyspnoi Hansen and Sørensen, 1904 (Arachnida: Opiliones)". ResearchGate. Retrieved 2019-05-24.
  2. "A new species of the genus Giljarovia Kratochvíl, 1959 (Arachnida:Opiliones: Phalangiidae) from the Crimea". ResearchGate. Retrieved 2019-05-23.
  3. "Hadzinia ferrani, sp. n. (Opiliones: Nemastomatidae), a highly specialized troglobiotic harvestman from Slovenia". ResearchGate. Retrieved 2019-05-23.
  4. "Mediostoma izmirica sp. nov.: A new harvestman species (Arachnida, Opiliones, Nemastomatidae) from Turkey". ResearchGate. Retrieved 2019-05-23.
  5. Tchemeris, A. N. (2013). "Two new harvestman species (Arachnida: Opiliones) from the collection of Siberian Zoological Museum". Arthropoda Selecta. 22: 41046–0. doi:10.15298/arthsel.22.1.03 – via ResearchGate.
  6. Karaman, IVO M. (2013). "Nemaspela ladae sp. N., a new troglobitic nemastomatid (Opiliones, Dyspnoi, Nemastomatidae) from a Dinaric cave". Zootaxa. 3694 (3): 240–248. doi:10.11646/zootaxa.3694.3.4. PMID 26312286 – via ResearchGate.
  7. Martens, Jochen (2016). "Sinostoma yunnanicum, the first nemastomatine harvestman in China (Arachnida: Opiliones: Nemastomatidae)". Zootaxa. 4126 (3): 444–450. doi:10.11646/zootaxa.4126.3.9. PMID 27395599 – via ResearchGate.
  8. Martens, Jochen (2017). "On Starengovia Snegovaya, a genus of Asian nemastomatines (Arachnida: Opiliones: Nemastomatidae)". Revue Suisse de Zoologie. 124 (2): 187–201. doi:10.5281/zenodo.893462. S2CID 52200517 – via ResearchGate.
  9. Zhang, Chao; Martens, Jochen (2018). "Ancient home or in exile? The easternmost species of genus Starengovia Snegovaya, 2010 found in China (Opiliones, Nemastomatidae, Nemastomatinae)". ZooKeys (770): 105–115. doi:10.3897/zookeys.770.25491. ISSN 1313-2970. PMC 6041357. PMID 30002591.
  10. "A summary list of fossil spiders and their relatives" (PDF). wsc.nmbe.ch. Retrieved 2019-05-23.
  11. "New fossil harvestmen (Arachnida: Opiliones) in the HOFFEINS ambe...: Ingenta Connect". www.ingentaconnect.com. Archived from the original on 2020-09-17.
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