Ulmus × hollandica

In form and foliage, the trees are broadly intermediate between the two species.[8] F1 hybrids between wych and field elm are fully fertile, but produce widely variant progeny.[1] Many also inherit the suckering habit of their field elm parent.[9] Both Richens and Rackham noted that examples in the East Anglian hybridization zone were sometimes pendulous in form.[1][10]

• 
  Ulmus × hollandica in Lincolnshire hedgerow
• 
  Nascent samarae of same

Some examples of the hybrid possess a moderate resistance to Dutch elm disease.[11]

The hybrid has been introduced to North America and Australasia.

The great elm in The Grove of Magdalen College, Oxford, photographed by Henry Taunt in 1900,[12] long believed to be a wych elm before being identified by Elwes as a 'Vegeta'-type hybrid, was for a time the largest elm known in Britain before it was blown down in 1911. It measured 44 m tall, its trunk at breast height being 2.6 m in diameter,[13] and comprised an estimated 81 cubic metres (2,900 cu ft) of timber,[14] making it the largest tree of any kind in Britain and possibly the largest north of the Alps.[15] However, as Elwes pointed out, its calculated age would place its planting in the late 17th or early 18th century,[16] long before the introduction of the Huntingdon elm, making the tree in question more likely to be a Chichester elm. A second tree nearby, described by Elwes as "similar in habit and foliage" (see 'External links') and 130 feet (40 m) tall by 23 feet (7.0 m) in girth in 1912,[17] was confirmed by Nellie Bancroft in a Gardener's Chronicle article in 1934 as a 'Vegeta'-type hybrid;[18] it was propagated by Heybroek in 1958 and cultivated at the Baarn elm research institute as clone P41.[19][note 1] The tree survived till the 1960s. Like the Queens' College Chichester elms in Cambridge, the Magdalen College trees were not observed to produce root suckers, though The Grove at Magdalen has long been a deer park, and any sucker growth is likely to have been cropped.[16] The Oxford zoologist Robert Gunther attributed the larger tree's unusual size to the fact (discovered in 1926) that it had been growing on a phosphate-rich bone-bed, made up of the remains of mammoths and other prehistoric animals.[20]

• 
  The great elm in The Grove, Magdalen College, Oxford (ca. 1870)[21]
• 
  Magdalen College elm in leaf (1899)[21][22]
• 
  Magdalen College elm (1906)
• 
  Magdalen College elm after being blown down (1911)[23]

With a girth of 6.9 m (22.6 ft) and a height of 40 metres (130 ft), the Ulmus × hollandica hybrid elm on Great Saling Green, Great Saling, near Braintree, Essex, reckoned at least 350 years old,[24] was reputedly the largest elm in England, before succumbing to Dutch Elm Disease in the 1980s;[25] Elwes and Henry (1913) misidentified it as U. nitens (Ulmus minor).[5]

Examples of mature survivors in the East Anglian hybridisation zone include those near Royston, Hertfordshire, designated 'Elm of the Year, 2004' by Das Ulmen Büro.[26] An example of the weeping form survives at Actons Farm, Sawbridgeworth, Hertfordshire.[27]

There are two notable TROBI Champion trees in the British Isles, one at Little Blakenham, Suffolk, measuring 160 centimetres (5.2 ft) d.b.h. in 2008, the other at Nounsley, Essex, 17 metres (56 ft) high by 150 centimetres (4.9 ft) d.b.h. in 2005.[28]

The so-called W. G. Grace Elm, a majestic spreading hybrid (labelled just "Dutch elm"), the last survivor (2009) of a ring of elms round East Oval, Ballarat, Victoria, was reputedly planted by W. G. Grace himself in 1874 during a tour by the England cricket team.[29] It has been shown, however, to date from 1900.[30] 'W. G. G.' had attained a spread round the middle of 31 m by 1982.[31]

At least 40 cultivars have been recorded, although some may not have survived Dutch elm disease:

Others provisionally identified as Ulmus × hollandica include 'Scampstoniensis' and 'Virens (by Green); 'Purpurea' and 'Louis van Houtte' (by F. J. Fontaine); 'Escaillard' and 'Hillieri' (by Buisman); 'Rugosa Pendula' (by Arnold and Morton arboreta); and Späth's 'Fastigiata Glabra' (by Melville). Cultivars at one time or another identified as U. × hollandica, but which may have suffered misidentification through confusion with U. glabra Huds. cultivars that share the same name include 'Fastigiata Glabra', 'Fastigiata Macrophylla', 'Latifolia', and 'Rugosa'. In the 19th century and early 20th, Ulmus × hollandica cultivars (as well as those of wych elm) were often grouped under Ulmus montana.[13][32][33]

• 
  An unidentified U. × hollandica cultivar (not 'Vegeta') on Leith Links, Edinburgh (fruiting, April 2017)
• 
  Samarae
• 
  July
• 
  Short shoot undersides
• 
  Short shoot
• 
  Bark

• 
  An unidentified smooth-leaved U. × hollandica cultivar, North Merchiston Cemetery, Edinburgh (2016)
• 
  Same, fruiting
• 
  Fruit
• 
  Foliage
• 
  Dried leaves, showing a longer petiole than that of wych elm
• 
  Bark

The elms in the Suffolk landscape-paintings and drawings of John Constable were "most probably East Anglian hybrid elms ... such as still grow in the same hedges" in Dedham Vale and East Bergholt.[34] (His Flatford Mill elms were U. minor.[35]) Elm trees in Old Hall Park, East Bergholt, showing a clump of these hybrids, is often considered the finest of Constable's elm-studies.[36][37]

• 
  John Constable, Stoke-by-Nayland, Suffolk, c.1830
• 
  John Constable, The Cornfield, 1826 (lane between East Bergholt and Dedham) [38]
• 
  John Constable, Elm trees in Old Hall Park, East Bergholt, 1817 (Ulmus × hollandica[36])

• "Herbarium specimen - L.1582475". Botany catalogues. Naturalis Biodiversity Center. Labelled U. hollandica aff. vegeta, Magdalen College Grove specimen 1 (Heybroek, Oxford 1958)
• "Herbarium specimen - L.1582476". Botany catalogues. Naturalis Biodiversity Center. Labelled U. hollandica aff. vegeta, Magdalen College Grove specimen 2 (Heybroek, Oxford 1958)