Synonymy

• Felis atrox Leidy, 1853: 321 [protonym]
• Felis imperialis Leidy, 1873: 260
• Felis atrox bebbi Merriam, 1909: 301
• Panthera leo atrox
• Panthera atrox

Primary references

• Leidy, J. 1853. Description of an extinct species of American Lion: Felis atrox. Transactions of the American Philosophical Society n.s. 10: 319–321. DOI: 10.2307/1005282 JSTOR BHL Reference page. 

• Merriam, J.C. 1909. The skull and dentition of an extinct cat closed allied to Felis atrox Leidy. University of California Publications, Bulletin of the Department of Geological Sciences 5(20): 291–304. HathiTrust Reference page. 

Additional references

• Merriam, J.C. & Stock, C. 1932. The Felidae of Rancho La Brea. Carnegie Institution of Washington. HathiTrust Reference page.  pp. 164–199.

• Kurtén, B. 1985. The Pleistocene lion of Beringia. Annales Zoologici Fennici 22: 117–121. PDF Reference page. 

• Barnett, R., Mendoza, M.L.Z., Soares, André Elias Rodrigues, Ho, S.Y.W., Zazula, G., Yamaguchi, N., Shapiro, B., Kirillova, I.V., Larson, G. & Gilbert, M.T.P. 2016. Mitogenomics of the Extinct Cave Lion, Panthera spelaea (Goldfuss, 1810), Resolve its Position within the Panthera Cats. Open Quaternary 2(4): 1–11. DOI: 10.5334/oq.24 ResearchGate Reference page.