2014 in arthropod paleontology

This list of fossil arthropods described in 2014 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2014. The list only includes taxa at the level of genus or species.

List of years in arthropod paleontology
In paleontology
2011
2012
2013
2014
2015
2016
2017
In science
2011
2012
2013
2014
2015
2016
2017
In reptile paleontology
2011
2012
2013
2014
2015
2016
2017
In paleobotany
2011
2012
2013
2014
2015
2016
2017
In paleoentomology
2011
2012
2013
2014
2015
2016
2017
In paleomalacology
2011
2012
2013
2014
2015
2016
2017
In archosaur paleontology
2011
2012
2013
2014
2015
2016
2017
In mammal paleontology
2011
2012
2013
2014
2015
2016
2017
In paleoichthyology
2011
2012
2013
2014
2015
2016
2017

Arachnids

Newly named taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Baalzebub? mesozoicum[1]

Sp. nov

Valid

Penney

Cretaceous (middle Cenomanian to early Santonian)

 France

A spider of uncertain placement. Originally described as a ray spider; Wunderlich & Müller (2020) transferred it to the family Zarqaraneidae.[2]

Cheirolepidoptus[3]

Gen. et sp. nov

Valid [4]

Sidorchuk & Lindquist in Sidorchuk et al.

Late Triassic (Carnian)

 Italy

A Tetrapodili mite
The type species is C. dolomiticus.

Feaella (Tetrafeaella) groehni[5]

Sp. nov

Valid

Henderickx in Henderickx & Boone

Eocene

Baltic amber

 Europe

A feaellid pseudoscorpion

Hastocularis[6]

Gen. et sp. nov

Valid

Garwood et al.

Carboniferous (late Stephanian)

 France

A Tetrophthalmi Opiliones.
The type species is H. argus.

Kronocharon[7]

Gen. et sp. nov

Valid

Engel & Grimaldi

Cretaceous (latest Albian or earliest Cenomanian)

Burmese amber

 Myanmar

An Amblypygi originally assigned to Neoamblypygi
excluded from this group by Garwood et al., 2017[8]). The type species is Kronocharon prendinii.

Mayachernes[9]

Gen. et sp. nov

Jr synonym

Riquelme et al.

Miocene

Mexican amber

 Mexico

A chernetid pseudoscorpion. The type species is Mayachernes maatiatus.
Moved to Byrsochernes maatiatus by Judson (2016)[10]

Minyacarus[3]

Gen. et sp. nov

Valid[4]

Sidorchuk & Lindquist in Sidorchuk et al.

Late Triassic (Carnian)

 Italy

A Tetrapodili mite.
The type species is M. aderces.

Montsecarachne[11]

Gen. et sp. nov

Valid

Selden

Early Cretaceous (c. 129 Ma)

El Montsec

 Spain

A plectreurid spider.
The type species is M. amicus.

?Opilioacarus groehni[12]

Sp. nov

Valid

Dunlop & Bernardi

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An opilioacarid mite.

Paracharonopsis[7]

Gen. et sp. nov

Valid

Engel & Grimaldi

Eocene (Ypresian)

Cambay Shale Formation

 India

A Paracharontidae Amblypygi.
The type species is P. cambayensis.

Paratrombium rovniense[13]

Sp. nov

Valid

Konikiewicz & Mąkol

Eocene

Rovno amber

 Ukraine

A mite.

Pleophrynus hawesi[14]

Sp. nov

Valid

Dunlop et al.

Carboniferous (Pennsylvanian)

Astrasado Formation

 United States

An eophrynid trigonotarbid.

Pulchellaranea[15]

Gen. et sp. nov

Valid

Poinar

Cenozoic

Dominican amber

 Dominican Republic

An orb-weaver spider.
The type species is P. pedunculata.

Seppo[16]

Gen. et sp. nov

Valid

Selden & Dunlop

Early Jurassic (Toarcian)

Ciechocinek Formation

 Germany

A probable palpimanoidean spider.
The type species is S. koponeni.

Tityus knodeli[17]

Sp. nov

Valid

Lourenço

Miocene

Mexican amber

 Mexico

A buthid scorpion.

Tuckerella fossilibus[18]

Sp. nov

Valid

Khaustov, Sergeyenko & Perkovsky

Eocene

Rovno amber

 Ukraine

A peacock mite.

Newly named crustaceans

Name Novelty Status Authors Age Unit Location Notes Images

Acanthaxius carinatus[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidae, a species of Acanthaxius.

Acanthochirana liburiaensis[20]

Sp. nov

Valid

Garassino, Audo, Charbonnier & Schweigert in Bravi et al.

Middle Jurassic (Bajocian/Bathonian)

 Italy

A member of Aegeridae, a species of Acanthochirana.

Acanthochirana triassica[21]

Sp. nov

Valid

Garassino, Schweigert & Muscio

Late Triassic (Norian)

Dolomia di Forni Formation

 Italy

A member of Aegeridae, a species of Acanthochirana.

Adamanteryon[22]

Gen. et sp. nov

Valid

Audo et al.

Middle Jurassic

La Voulte-sur-Rhône Lagerstätte

 France

A member of Polychelida. The type species is Adamanteryon fourneti.

Alavatanais margulisae[23]

Sp. nov

Valid[24]

Sánchez-García, Peñalver & Delclòs in Sánchez-García et al.

Early Cretaceous

 Spain

A paratanaoid tanaid, a species of Alavatanais.

Alessandranina[25]

Gen. et comb. nov

Valid

Karasawa et al.

Late Cretaceous (Cenomanian)

 United Kingdom

A palaeocorystid crab. The type species is "Notopocorystes" (Cretacoranina) ornatus Wright and Collins (1972).

Ammopylocheles petersi[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Ammopylocheles.

Anisaeger[27]

Gen. et 2 sp. nov

Valid

Schweitzer et al.

Middle Triassic (Anisian)

 China

A member of Aegeridae. The type species is Anisaeger brevirostrus; genus also contains Anisaeger spiniferus.

Annuntidiogenes jurassicus[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Annuntidiogenes.

Antrimpos colettoi[21]

Sp. nov

Valid

Garassino, Schweigert & Muscio

Late Triassic (Norian)

Dolomia di Forni Formation

 Italy

A member of Penaeidae. Originally described as a species of Antrimpos, but subsequently transferred to the genus Ladinicaris.[28]

Araripenaeus[29]

Gen. et sp. nov

Valid

Pinheiro, Saraiva & Santana

Early Cretaceous (Albian)

Romualdo Formation

 Brazil

A member of Penaeoidea of uncertain phylogenetic placement. The type species is Araripenaeus timidus.

Arcania kobayashinobuakii[30]

Sp. nov

Valid

Karasawa

Middle Pleistocene

Toyohashi Formation

 Japan

A member of Leucosiidae, a species of Arcania.

Archaeoniscus italiensis[31]

Sp. nov

Valid

Jones, Feldmann & Garassino

Jurassic

 Italy

An isopod, a species of Archaeoniscus.

Ariecornibus[32]

Gen. et sp. nov

Valid

Franţescu

Early Cretaceous (late Albian)

 United States

A crab. The type species is Ariecornibus schweitzerae.

Axiopsis pawpawensis[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidae, a species of Axiopsis.

Axiopsis sampsonumae[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidae, a species of Axiopsis.

Axiopsis spinifera[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidae, a species of Axiopsis.

Bellcarcinus[33]

Gen. et sp. nov

Valid

Luque

Early Cretaceous (late Aptian)

Paja Formation

 Colombia

A crab belonging to the group Raninoidia, possibly a member of the family Orithopsidae. The type species is Bellcarcinus aptiensis.

Belos[32]

Gen. et sp. nov

Valid

Franţescu

Early Cretaceous (late Albian)

 United States

A crab. The type species is Belos trispinae.

Bolbozoe beccata[34]

Sp. nov

Valid[35]

Perrier et al.

Early Silurian

 Australia

A myodocope ostracod, a species of Bolbozoe.

Branchiocarcinus pacificus[36]

Sp. nov

Valid

Nyborg, Ossó & Vega

Late Cretaceous (Maastrichtian)

Moreno Formation

 United States

A portunoid crab. Originally described as a member of the family Icriocarcinidae and a species of Branchiocarcinus; Kornecki, Feldmann & Schweitzer (2017) considered the family Icriocarcinidae to be a junior synonym of the family Lithophylacidae and transferred B. pacificus to the genus Lithophylax.[37]

Brazilestheria[38]

Gen. et comb. nov

Valid

Chen & Shen

Early Cretaceous

 Brazil

A clam shrimp belonging to the family Sinoestheriidae. Genus includes "Asmussia" souzae Guerin-Franiatte & Taquet (1993).

Canalarta[34]

Gen. et sp. nov

Valid[35]

Perrier et al.

Early Silurian

 Australia

A myodocope ostracod. The type species is Canalarta papata.

Candonopsis alagoensis[39]

Sp. nov

Valid

Tomé, Lima Filho & Neumann

Early Cretaceous

Araripe basin

 Brazil

An ostracod.

Cantabroxanthus[40]

Gen. et sp. nov

Valid

Ossó & Díaz Isa

Late Cretaceous (middle Campanian)

 Spain

A feldmanniid crab, a relative of etyiids. The type species is Cantabroxanthus loredoensis.

Casertanus[20]

Gen. et sp. nov

Valid

Garassino, Audo, Charbonnier & Schweigert in Bravi et al.

Middle Jurassic (Bajocian/Bathonian)

 Italy

A member of Sergestidae. The type species is Casertanus sabellicus.

Ceratobairdia crassispinosa[41]

Sp. nov

Valid

Monostori & Tóth

Triassic

 Hungary

An ostracod, a species of Ceratobairdia.

Claudioranina[25]

Gen. et comb. nov

Valid

Karasawa et al.

Eocene (Lutetian-Bartonian)

 Italy

A raninid crab. The type species is "Cyrtorhina" oblonga Beschin, Busulini, De Angeli & Tessier (1988).

Clinocypris epacrus[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Clinocypris.

Colombicarcinus[25]

Gen. et sp. nov

Valid

Karasawa et al.

Early Cretaceous (Aptian)

 Colombia

A necrocarcinid crab. The type species is Colombicarcinus laevis.

Concinnalepas[43]

Gen. et comb. nov

Valid

Gale

Late Jurassic

 United Kingdom

A zeugmatolepadid barnacle. A new genus for "Pollicipes" concinnus Morris (1845).

Cophinia grekoffi[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Cophinia.

Cophinia ovalis[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Cophinia.

Corallomursia[46]

Gen. et sp. nov

Valid

De Angeli & Ceccon

Early Eocene

 Italy

A crab belonging to the family Calappidae. Genus includes new species C. eocaena.

Ctenocheles fritschi[47]

Sp. nov

Valid

Hýžný, Kočová Veselská & Dvořák

Late Cretaceous (Coniacian)

Březno Formation

 Czech Republic

A ctenochelid axiidean, a species of Ctenocheles.

Cycleryon romani[22]

Sp. nov

Valid

Audo et al.

Middle Jurassic

La Voulte-sur-Rhône Lagerstätte

 France

A member of Polychelida, a species of Cycleryon.

Cymothoidana[48]

Gen. et sp. nov

Valid

Jarzembowski et al.

Early Cretaceous

Wealden Supergroup

 United Kingdom

A cymothoidan isopod. The type species is Cymothoidana websteri.

Cypria poietes[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Cypria.

Cytheropteron paramassoni[49]

Sp. nov

Valid

Yasuhara & Okahashi

Late Quaternary

North Atlantic Ocean (Rockall Plateau)

A cytherurid podocopid ostracod, a species of Cytheropteron.

Damonella grandiensis[39]

Sp. nov

Valid

Tomé, Lima Filho & Neumann

Early Cretaceous

Araripe basin

 Brazil

An ostracod.

Dawsonius tigris[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidea belonging to the family Ctenochelidae, a species of Dawsonius.

Devonostenopus[50]

Gen. et sp. nov

Valid

Jones et al.

Devonian

Huntley Mountain Formation

 United States

A member of Stenopodidea. The type species is Devonostenopus pennsylvaniensis.

Dicerobairdia latispinosa[41]

Sp. nov

Valid

Monostori & Tóth

Triassic

 Hungary

An ostracod, a species of Dicerobairdia.

Diogenicheles[51]

gen et sp nov

Valid

Fraaije et al.

Oxfordian

 Poland

A parapylochelid hermit crab. The type species is Diogenicheles theodorae.

Distaeger[27]

Gen. et sp. nov

Valid

Schweitzer et al.

Middle Triassic (Anisian)

 China

A member of Aegeridae. The type species is Distaeger prodigiosus.

Distefania(?) tangishirazensis[52]

Sp. nov

Valid

Jagt et al.

Early Cretaceous (Albian)

Kazhdumi Formation

 Iran

A member of Dromioidea, possibly a species of Distefania.

Distefania(?) vanrijsselti[53]

Sp. nov

Valid

Jagt, Fraaije & van Bakel

Late Cretaceous (late Maastrichtian)

Maastricht Formation

 Netherlands

A member of Dromioidea, possibly a species of Distefania.

Dromia evae[54]

Sp. nov

Valid

Collins

Miocene

 Austria

A member of Dromiidae, a species of Dromia.

Dromiopsis paleogenicus[46]

Sp. nov

Valid

De Angeli & Ceccon

Early Eocene

 Italy

A crab belonging to the family Dynomenidae.

Dundgobiestheria[55]

Gen. et sp. nov

Valid

Li in Li et al.

Probably Middle Jurassic

Eedemt Formation

 Mongolia

A clam shrimp. The type species is Dundgobiestheria mandalgobiensis.

Electrotanais[23]

Gen. et sp. nov

Valid[24]

Sánchez-García, Peñalver & Delclòs in Sánchez-García et al.

Early Cretaceous

 Spain

A paratanaoid tanaid. The type species is Electrotanais monolithus.

Entomozoe bispinosa[56]

Sp. nov

Valid

Wang & Wu

Early Silurian

Shimenkan Formation

 China

An ostracod, a species of Entomozoe.

Entomozoe suboblonga[56]

Sp. nov

Valid

Wang & Wu

Early Silurian

Shimenkan Formation

 China

An ostracod, a species of Entomozoe.

Entomozoe wukeensis[56]

Sp. nov

Valid

Wang & Wu

Early Silurian

Shimenkan Formation

 China

An ostracod, a species of Entomozoe.

Eogordonella[57]

Gen. et sp. nov

Valid

Garassino et al.

Eocene

 Iran

A solenocerid penaeoid. The type species is Eogordonella iranianiensis.

Eomaldivia? friebei[54]

Sp. nov

Valid

Collins

Miocene

 Austria

A member of Trapeziidae, possibly a species of Eomaldivia.

Eonatatolana[58]

Gen. et sp. nov

Valid[59]

Etter

Middle Jurassic

 Germany

A cirolanid isopod. The type species is Eonatatolana geisingensis.

Eopabdehus[57]

Gen. et sp. nov

Valid

Garassino et al.

Eocene

 Iran

A penaeid. The type species is Eopabdehus babaheydariensis.

Eosestheria lushangfenensis[60]

Sp. nov

Valid

Liao, Shen & Huang

Early Cretaceous

Lushangfen Formation

 China

A clam shrimp, a species of Eosestheria.

Epibrachylepas[61]

Gen. et sp. et comb. nov

Valid[62]

Gale in Gale & Sørensen

Late Cretaceous (Campanian and Maastrichtian)

 Netherlands
 Sweden

A basal balanomorph barnacle. The type species is Epibrachylepas newmani; genus also includes Epibrachylepas smeetsi (Bosquet, 1857).

Eryma osciensis[20]

Sp. nov

Valid

Garassino, Audo, Charbonnier & Schweigert in Bravi et al.

Middle Jurassic (Bajocian/Bathonian)

 Italy

A member of the family Erymidae.

Etcheslepas[43]

Gen. et sp. et comb. nov

Valid

Gale

Late Jurassic

 United Kingdom

A member of Pollicipedidae. Genus contains the new species Etcheslepas durotrigensis, as well as "Zeugmatolepas" fragilis Withers (1928).

Etisus evamuellerae[63]

Sp. nov

Valid

Hyžný, van Bakel & Guinot

Middle Miocene

 Austria
 Hungary
 Spain?

A member of Xanthidae, a species of Etisus.

Eucytherura myrsinae[64]

Sp. nov

Valid

Uffenorde in Uffenorde & Radtke

Oligocene

 Germany

A cytherurid podocopid ostracod.

Eucytherura zehali[49]

Nom. nov

Valid

Yasuhara & Okahashi

Late Quaternary

North Atlantic Ocean (Carolina Slope)

A cytherurid podocopid ostracod; a replacement name for Eucytherura hazeli Yasuhara, Okahashi & Cronin (2009).

Eukloedenella adcapitisdolorella[65]

Sp. nov

Valid[66]

Forel

Early Triassic

Kokarkuyu Formation

 Turkey

An ostracod, a species of Eukloedenella.

Eurotanais[23]

Gen. et sp. nov

Valid[24]

Sánchez-García, Peñalver & Delclòs in Sánchez-García et al.

Early Cretaceous

 Spain

A paratanaoid tanaid. The type species is Eurotanais terminator.

Falklandicypris[42]

Gen. et 2 sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod. The type species is Falklandicypris petrasaltata; genus also contains Falklandicypris desiderata.

Faxelepas[67]

Gen. et comb. nov

Valid[68]

Gale

Late Cretaceous (Maastrichtian) to late Miocene[69]

 Denmark
 Italy
 Netherlands
 Spain[69]
 Sweden

A barnacle related to verrucomorphs. A new genus for "Pycnolepas" bruennichi Withers (1914); genus also includes "Pycnolepas" paronai De Alessandri (1895).[69]

Fossocytheridea potiguarensis[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Fossocytheridea.

Fossocytheridea tiberti[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Fossocytheridea.

Gangamoncythere[42]

Gen. et sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod. The type species is Gangamoncythere colini.

Glabropilumnus nitidus[54]

Sp. nov

Valid

Collins

Miocene

 Austria

A member of Pilumnidae, a species of Glabropilumnus.

Globotalicypridea mirabilis[70]

Sp. nov

Valid

Silye, Colin & Codrea

Late Cretaceous (Maastrichtian)

 Romania

A talicyprideine cypridid ostracod, a species of Globotalicypridea.

Gracilimanus[71]

Gen. et sp. nov

Valid

Feldmann & Schweitzer in Guyer, Schweitzer & Feldmann

Middle Jurassic (Bajocian/Bathonian)

Gypsum Springs Formation

 United States

A member of Astacidea, possibly a member of the family Stenochiridae. The type species is Gracilimanus obscurus.

Giulianolyreidus[25]

Gen. et comb. nov

Valid

Karasawa et al.

Paleocene (Thanetian)

 United States

A lyreidid crab. The type species is "Symethis" johnsoni Rathbun (1935); genus also contains "Symnista" bidentata Rathbun (1935).

Gypsonicus[71]

Gen. et sp. nov

Valid

Schweitzer & Feldmann in Guyer, Schweitzer & Feldmann

Middle Jurassic (Bajocian/Bathonian)

Gypsum Springs Formation

 United States

A member of Astacidea belonging to the family Stenochiridae. The type species is Gypsonicus wyomingensis.

Harpactocarcinus fedrigoi[72]

Sp. nov

Valid

Beschin, De Angeli & Zorzin

Eocene

 Italy

A zanthopsid carpilioid crab, a species of Harpactocarcinus.

Haughtonileberis dinglei[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian to Santonian)

 Brazil
 Nigeria

A trachyleberidid ostracod, a species of Haughtonileberis.

Hemicytherura viviersae[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherurid ostracod, a species of Hemicytherura.

Hepatus pauli[73]

Sp. nov

Valid

Collins, Garvie & Mellish

Pleistocene

Beaumont Formation

 United States

A member of Aethridae, a species of Hepatus.

Hourcqia woodi[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Hourcqia.

Housacheles ovalis[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Housacheles.

Hyas chippubetsuensis[74]

Sp. nov

Valid

Kato, Nakashima & Yanagisawa

Early Pliocene

Chippubetsu Formation

 Japan

A member of Oregoniidae, a species of Hyas.

Hyas tentokujiensis[74]

Sp. nov

Valid

Kato, Nakashima & Yanagisawa

Early Pliocene

Tentokuji Formation

 Japan

A member of Oregoniidae, a species of Hyas.

Icenilepas[43]

Gen. et sp. nov

Valid

Gale

Late Cretaceous (Santonian)

 United Kingdom

A zeugmatolepadid barnacle. The type species is Icenilepas collinsi.

Ilhasina? leiodermatus[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, possibly a species of Ilhasina.

Ilyocypris bisulcata[75]

Sp. nov

Valid

Qu et al.

Early Paleocene

Mingshui Formation

 China

An ilyocypridid cypridoid ostracod, a species of Ilyocypris.

Italianira[25]

Gen. et comb. nov

Valid

Karasawa et al.

Eocene (Ypresian)

 Italy

A raninid crab. The type species is "Ranilia" punctulata Beschin, Busulini, De Angeli & Tessier (1988).

Jandairella[44]

Gen. et sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A trachyleberidid ostracod. The type species is Jandairella obesa.

Kegelina[76]

Gen. et comb. nov

Valid

de Queiroz Neto, Sames & Colin

Early Cretaceous

 Brazil
 Gabon
 United States

A cypridoid ostracod. A new genus for "Cypridea" anomala Peck (1941), "Cypridea" armata Krömmelbein (1962), "Cypridea" bisculpturata Wicher (1959), "Cypridea" depressa Moura (1972) and "Cypridea" kegeli Wicher (1959).

Kotoracythere paiki[77]

Sp. nov

Valid

Lee

Pliocene-Pleistocene

Seogwipo Formation

 South Korea

An ostracod, a species of Kotoracythere.

Krausella danukalovae[78]

Sp. nov

Valid

Melnikova & Danukalova

Ordovician

 Russia

An ostracod, a species of Krausella.

Kuzminaella arabieni[79]

Sp. nov

Valid

Groos-Uffenorde in Groos-Uffenorde & Rabien

Devonian

 Germany

An ostracod.

Lepidophthalmus paratethyensis[80]

Sp. nov

Valid[81]

Gašparič & Hyžný

Early Miocene (late Burdigalian)

 Slovenia

A member of Callianassidae, a species of Lepidophthalmus.

Longiscula kazarkina[78]

Sp. nov

Valid

Melnikova & Danukalova

Ordovician

 Russia

An ostracod, a species of Longiscula.

Looneyellopsis tuberculatus[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Looneyellopsis.

Loxocorniculum? narendrai[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A loxoconchid ostracod, possibly a species of Loxocorniculum.

Luprisca[82]

Gen. et sp. nov

Valid

Siveter et al.

Ordovician (Katian)

Lorraine Group

 United States

A myodocope ostracod. The type species is Luprisca incuba.

Martillepas[43]

Gen. et comb. nov

Valid

Gale

Late Jurassic

 United Kingdom

A zeugmatolepadid barnacle. A new genus for "Calantica" ovalis Withers (1928), "Zeugmatolepas" costata Withers (1928) and "Zeugmatolepas" hollisi Withers (1928).

Masticacheles[51]

Gen et 2 sp nov

Valid

Fraaije et al.

Oxfordian to Kimmeridgian

 Germany
 Poland

A parapylochelid hermit crab. The type species is Masticacheles longirostris from Oxfordian of Poland;[51] subsequently a second species Masticacheles minimus from late Kimmeridgian of Germany was named by Fraaije (2014).[26]

Mesoparapylocheles jaegeri[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Mesoparapylocheles.

Mesoparapylocheles schweigerti[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Mesoparapylocheles.

Monoceratina hussonae[65]

Sp. nov

Valid[66]

Forel

Early Triassic

Kokarkuyu Formation

 Turkey

An ostracod, a species of Monoceratina.

Montemagrellus[46]

Gen. et sp. nov

Valid

De Angeli & Ceccon

Early Eocene

 Italy

A crab belonging to the family Crossotonotidae. Genus includes new species M. denticulatus.

Muelleristhes[83]

Gen. et comb. nov

Valid

Garassino, De Angeli & Pasini

Late Cretaceous (Cenomanian-Turonian)

 Morocco

A porcelain crab; a new genus for "Paragalathea" africana Garassino, De Angeli & Pasini (2008).

Munidopsis palmuelleri[84]

Sp. nov

Valid

Hyžný et al.

Middle Miocene

Slovenske Gorice Formation

 Slovenia

A munidopsid galatheoid, a species of Munidopsis.

Musacchiocythere[42]

Gen. et comb. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod. The type species is Musacchiocythere sarunata (Musacchio, 1978).

Neopilumnoplax pohorjensis[80]

Sp. nov

Valid[81]

Gašparič & Hyžný

Early Miocene (late Burdigalian)

 Slovenia

A member of Mathildellidae, a species of Neopilumnoplax.

Nothocarapacestheria[85]

Gen. et sp. nov

Valid

Rohn, Dutra & Cabral

Probably Jurassic

Caturrita Formation

 Brazil

A clam shrimp belonging to the family Eosestheriidae. The type species is Nothocarapacestheria soturnensis.

Occultocaris[86]

Gen. et sp. nov

Valid

Winkler

Late Jurassic (early Tithonian)

Solnhofen Plattenkalk

 Germany

A member of Caridea, possibly a bresilioid. The type species is Occultocaris frattigianii.

Ovocytheridea anterocompressa[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Ovocytheridea.

Ovocytheridea posteroprojecta[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Ovocytheridea.

Ovocytheridea reymenti[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous

 Brazil
 Nigeria

A cytherideid ostracod, a species of Ovocytheridea.

Ovocytheridea triangularis[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Ovocytheridea.

Pagurus retznensis[54]

Sp. nov

Valid

Collins

Miocene

 Austria

A hermit crab, a species of Pagurus.

Pagurus texensis[32]

Sp. nov

Valid

Franţescu

Early Cretaceous (late Albian)

 United States

A hermit crab, a species of Pagurus.

Palaega furcillanatis[31]

Sp. nov

Valid

Jones, Feldmann & Garassino

Jurassic

 Italy

An isopod, a species of Palaega.

Palaeogammarus debroyeri[87]

Sp. nov

Valid

Jażdżewski, Grabowski & Kupryjanowicz

Eocene

Probably  Russia (Kaliningrad Oblast)

A member of Amphipoda found in Baltic amber, a species of Palaeogammarus.

Palinurellus bericus[88]

Sp. nov

Valid

De Angeli & Garassino

Eocene (Priabonian)

 Italy

A furry lobster, a species of Palinurellus.

Palladiocarcinus[46]

Gen. et sp. nov

Valid

De Angeli & Ceccon

Early Eocene

 Italy

A crab belonging to the family Pilumnidae. Genus includes new species P. brevidentatus.

Palmunidopsis[89]

Gen. et sp. nov

Valid

Fraaije

Miocene (Tortonian)

Pakhna Formation

 Cyprus

A munidopsid galatheoid. The type species is Palmunidopsis muelleri.

Parabrachylepas[61]

Gen. et comb. nov

Valid[62]

Gale in Gale & Sørensen

Late Cretaceous (Campanian)

 Sweden

A basal balanomorph barnacle. The type species is Parabrachylepas ifoensis (Withers, 1935).

Paracypris ? krivipotokensis[90]

Sp. nov

Valid

Forel in Sudar et al.

Early Triassic (Olenekian)

Obnica Formation

 Serbia

A cypridoid ostracod belonging to the family Paracyprididae, possibly a species of Paracypris.

Paracypris vietnamensis[91]

Sp nov

Valid

Tanaka & Komatsu in Shigeta et al.

Early Triassic (Olenekian)

Bac Thuy Formation

 Vietnam

An ostracod, a species of Paracypris.

Paraplesiocypridea[42]

Gen. et sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod. The type species is Paraplesiocypridea alloios.

Parapylochelitergites deductius[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Parapylochelitergites.

Parapylochelitergites prepustulosus[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Parapylochelitergites.

Paraxiopsis erugatus[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidae, a species of Paraxiopsis.

Parsacus[57]

Gen. et sp. et comb. nov

Valid

Garassino et al.

Eocene

 Iran
 Italy

A slipper lobster. Genus contains new species Parsacus eocenicus, as well as "Parribacus" cristatus Förster (1984).

Pauline nivisis[92]

Sp. nov

Valid

Perrier et al.

Silurian (late Telychian)

Pentamerus Bjerge Formation

 Greenland

A cylindroleberidid myodocope ostracod, a species of Pauline.

Paulinecaris[93]

Gen. et sp. nov

Valid[94]

Waloszek et al.

Cambrian

 Sweden

An entomostracan of uncertain phylogenetic placement. The type species is Paulinecaris siveterae.

Pedupycnolepas[67]

Gen. et comb. nov

Valid[68]

Gale

Early Cretaceous (probably early Aptian)

Antarctica (Alexander Island)

A barnacle; a new genus for "Pycnolepas" articulata Collins (1980).

Periacanthus tetracornis[95]

Sp. nov

Valid

Ferratges et al.

Eocene

 Spain

A crab belonging to the family Epialtidae.

Perissocytheridea caudata[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Perissocytheridea.

Perissocytheridea jandairensis[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Perissocytheridea.

Perissocytheridea mossoroensis[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherideid ostracod, a species of Perissocytheridea.

Petrochirus sanctilazzari[96]

Sp. nov

Valid

Baldanza et al.

Early Pleistocene

 Italy

A diogenid hermit crab, a species of Petrochirus.

Petrolisthes albianicus[32]

Sp. nov

Valid

Franţescu

Early Cretaceous (late Albian)

 United States

Originally classified as a porcelain crab belonging to the genus Petrolisthes. Robins & Klompmaker (2019) transferred it to the genus Hispanigalathea belonging to the galatheoid family Catillogalatheidae.[97]

Pilgrimcheles[51]

gen et 3 sp nov

Valid

Fraaije et al.

Oxfordian to Kimmeridgian

 Germany
 Poland

A parapylochelid hermit crab. The type species is Pilgrimcheles karolinae from Oxfordian of Poland;[51] subsequently additional two species, Pilgrimcheles kersteni and Pilgrimcheles vonmeyeri, were described from Kimmeridgian of Germany by Fraaije (2014).[26]

Pisidia? subnodosa[54]

Sp. nov

Valid

Collins

Miocene

 Austria

A porcelain crab, possibly a species of Pisidia.

Planobranchia palmuelleri[98]

Sp. nov

Valid

Artal, van Bakel & Onetti

Eocene (Lutetian)

Coll de Malla Formation

 Spain

A member of Inachidae, a species of Planobranchia.

Plioaxius texensis[19]

Sp. nov

Valid

Frantescu

Early Cretaceous (Albian)

Paw Paw Formation

 United States

A member of Axiidae, a species of Plioaxius.

Portunus muelleri[54]

Sp. nov

Valid

Collins

Miocene

 Austria

A member of Portunidae, a species of Portunus.

Potiguarella[44]

Gen. et 2 sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

 Brazil
 Gabon

A trachyleberidid ostracod. The type species is Potiguarella grosdidieri; genus also contains Potiguarella coimbrai.

Priscoverruca[67]

Gen. et comb. et sp. nov

Valid[68]

Gale

Late Cretaceous

 Belgium
 Germany
 Netherlands
 United Kingdom

A barnacle. The type species is "Verruca" prisca Bosquet (1854); genus also includes new species P. elongata.

Procytherura ballentae[44]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Turonian)

Potiguar Basin

 Brazil

A cytherurid ostracod, a species of Procytherura.

Protocosta babinoti[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A trachyleberidid ostracod, a species of Protocosta.

Proverruca dentifer[67]

Sp. nov

Valid[68]

Gale

Late Cretaceous (Campanian)

 Belgium
 Sweden

A barnacle.

Pylochelitergites exspectatus[99]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Oxfordian)

 Poland

A member of Pylochelidae, a species of Pylochelitergites.

Pyreneplax[100]

Gen. et sp. et comb. nov

Valid

Ossó, Domínguez & Artal

Eocene

 Italy
 Spain
 United States

A member of Vultocinidae. The type species is P. basaensis; genus also includes P. ranosa (Beschin, Busulini, De Angeli & Tessier, 2002), P. sandersi (Blow & Manning, 1997) and P. sommarugai (Beschin, Busulini & Tessier, 2009).

Quadratoplanus[32]

Gen. et sp. nov

Valid

Franţescu

Early Cretaceous (late Albian)

 United States

A crab. The type species is Quadratoplanus primitivus.

Retropluma slovenica[80]

Sp. nov

Valid[81]

Gašparič & Hyžný

Early Miocene (late Burdigalian)

 Slovenia

A member of Retroplumidae, a species of Retropluma.

Reviya sylvieae[65]

Sp. nov

Valid[66]

Forel

Early Triassic

Kokarkuyu Formation

 Turkey

An ostracod, a species of Reviya.

Schramidontus[101]

Gen. et sp. nov

Valid

Gueriau, Charbonnier & Clément

Late Devonian (Famennian)

 Belgium

A member of Eucarida related to Angustidontus. The type species is Schramidontus labasensis.

Scututergites[26]

Gen. et sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab. The type species is Scututergites anteroindentatus.

Semicytherura musacchioi[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A cytherurid ostracod, a species of Semicytherura.

Sphaeroma montefallanoense[31]

Sp. nov

Valid

Jones, Feldmann & Garassino

Jurassic

 Italy

An isopod, a species of Sphaeroma.

Stagmacaris subcircularis[26]

Sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab, a species of Stagmacaris.

Stevea martini[102]

Sp. nov

Valid

Feldmann, Schweitzer & Portell

Early Paleocene

Clayton Formation

 United States

A member of Hexapodidae, a species of Stevea.

Synurella aliciae[87]

Sp. nov

Valid

Jażdżewski, Grabowski & Kupryjanowicz

Eocene

Probably  Russia (Kaliningrad Oblast)

A member of Amphipoda found in Baltic amber, a species of Synurella.

Tamnacythere[77]

Gen. et sp. nov

Valid

Lee

Pliocene-Pleistocene

Seogwipo Formation

 South Korea

An ostracod. The type species is Tamnacythere seogwipoensis.

Tanycythere[103]

Gen. et 6 sp. nov

Valid

Cabral et al.

Early Jurassic to Early Cretaceous (Hettangian to Barremian?)

Western and central Europe

An ostracod. Genus contains six species: T. caudata, T. praecaudata (with two subspecies: T. praecaudata praecaudata and T. praecaudata parallela), T. duartei, T. posteroelongata, T. wattonensis and T. procera.

Tealliocaris walloniensis[104]

Sp. nov

Valid[105]

Gueriau, Charbonnier & Clément

Late Devonian (Famennian)

 Belgium

A pygocephalomorph crustacean,[106] a species of Tealliocaris.

Telamonocarcinus antiquus[107]

Sp. nov

Valid[108]

Luque

Early Cretaceous (early Albian)

 Colombia

A crab, a species of Telamonocarcinus.

Tesseropora canariana[109]

Sp. nov

Valid

Hornung

Pliocene (Zanclean)

 Spain ( Canary Islands)

A tetraclitid balanomorph barnacle, a species of Tesseropora.

Tethyseryon[20]

Gen. et sp. nov

Valid

Garassino, Audo, Charbonnier & Schweigert in Bravi et al.

Middle Jurassic (Bajocian/Bathonian)

 Italy

A member of Eryonidae. The type species is Tethyseryon campanicus.

Theriosynoecum ballentae[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Theriosynoecum.

Theriosynoecum petasmathylacus[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Theriosynoecum.

Tianzhuestheria jianchangensis[110]

Sp. nov

Valid

Wang

Middle–Late Jurassic

 China

A clam shrimp.

Timiriasevia fluitans[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Timiriasevia.

Triebelina anterotuberculata[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A bairdiid ostracod, a species of Triebelina.

Triebelina obliquocostata[45]

Sp. nov

Valid

Piovesan, Carbal & Colin in Piovesan et al.

Late Cretaceous (Santonian–Campanian)

Potiguar Basin

 Brazil

A bairdiid ostracod, a species of Triebelina.

Triglypta eedemtensis[55]

Sp. nov

Valid

Li in Li et al.

Probably Middle Jurassic

Eedemt Formation

 Mongolia

A clam shrimp, a species of Triglypta.

Triglypta yabraiensis[110]

Sp. nov

Valid

Wang

 China

A clam shrimp.

Ululapagurus[26]

Gen. et sp. nov

Valid

Fraaije

Late Jurassic (late Kimmeridgian)

 Germany

A hermit crab. The type species is Ululapagurus vanbakeli.

Unzhiella navis[111]

Sp. nov

Valid

Voronkova

Late Permian

 Russia

A volganellid ostracod, a species of Unzhiella.

Vecticypris samesi[42]

Sp. nov

Valid

Ayress & Whatley

Early Cretaceous

South Atlantic Ocean (North Falkland Basin)

A non-marine ostracod, a species of Vecticypris.

Verruca jagti[67]

Sp. nov

Valid[68]

Gale

Late Cretaceous (Maastrichtian)

Maastricht Formation

 Netherlands

A barnacle, a species of Verruca.

Volganella extensis[111]

Sp. nov

Valid

Voronkova

Late Permian

 Russia

A volganellid ostracod, a species of Volganella.

Volganella golubevi[111]

Sp. nov

Valid

Voronkova

Late Permian

 Russia

A volganellid ostracod, a species of Volganella.

Volganella vjaznikovensis[111]

Sp. nov

Valid

Voronkova

Late Permian

 Russia

A volganellid ostracod, a species of Volganella.

Voulteryon[22]

Gen. et sp. nov

Valid

Audo et al.

Middle Jurassic

La Voulte-sur-Rhône Lagerstätte

 France

A member of Polychelida. The type species is Voulteryon parvulus.

Wanestheria[38]

Gen. et comb. nov

Valid

Chen & Shen

Late Cretaceous

 China

A clam shrimp belonging to the family Sinoestheriidae. Genus includes "Sinoestheria" anhuiensis Chen & Shen (1982), "Sinoestheria" producta Chen & Shen (1982) and "Sinoestheria" semiorbita Chen & Shen (1982).

Youngiverruca[67]

Gen. et sp. nov

Valid[68]

Gale

Late Cretaceous (Maastrichtian)

 Germany

A barnacle. The type species is Youngiverruca ruegenensis.

Zovocarcinus[112]

Gen. et sp. nov

Valid

De Angeli & Garassino

Eocene (late Ypresian)

 Italy

A member of Panopeidae. The type species is Zovocarcinus muelleri.

Insects

Trilobites

Name Novelty Status Authors Age Unit Location Notes Images

Airophrys[113]

Gen. et sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae. The type species is A. balia.

Amphoton zangi[114]

Sp. nov

Valid

Sun, Jago & Bentley

Cambrian (Drumian)

Warburton Basin

 Australia

A polymerid trilobite, a species of Amphoton.

Anataphrus kermiti[115]

Sp. nov

Valid

Amati

Late Ordovician

Viola Springs Formation
Welling Formation

 United States

An isoteline asaphid, a species of Anataphrus.

Anataphrus megalophrys[115]

Sp. nov

Valid

Amati

Late Ordovician

Viola Springs Formation

 United States

An isoteline asaphid, a species of Anataphrus.

Asaphellus nazarenensis[116]

Sp. nov

Valid

Tortello & Esteban

Early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Asaphellus.

Asthenopsis conandersoni[117]

Sp. nov

Valid

Bentley & Jago

Cambrian

 Australia

Astroproetus franklini[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae.

Bathyurina curtisi[118]

Sp. nov

Valid

Adrain, Karim & Westrop

Ordovician (Floian)

Fillmore Formation

 United States

A bathyurid.

Bathyurina hooki[118]

Sp. nov

Valid

Adrain, Karim & Westrop

Ordovician (Floian)

Fillmore Formation

 United States

A bathyurid.

Bathyurina morrisi[118]

Sp. nov

Valid

Adrain, Karim & Westrop

Ordovician (Floian)

Fillmore Formation

 United States

A bathyurid.

Bathyurina sumneri[118]

Sp. nov

Valid

Adrain, Karim & Westrop

Ordovician (Floian)

Fillmore Formation

 United States

A bathyurid.

Bristolia colberti[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Bristolia.

Cambrunicornia saxonica[120]

Sp. nov

Valid

Geyer, Buschmann & Elicki

Middle Cambrian

Tröbitz Formation

 Germany

A species of Cambrunicornia.

Caudillaenus[121]

Gen. et sp. nov

Valid

Rábano, Gutiérrez-Marco & García-Bellido

Ordovician (late Darriwilian)

Taddrist Formation

 Morocco

An illaenid. The type species is Caudillaenus nicolasi.

Chapmanopyge knudseni[122]

Sp. nov

Valid

McCobb et al.

Early Ordovician

 Greenland

A species of Chapmanopyge.

Cheirurus falcatus[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Cheiruridae.

Cybantyx nebulosus[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae.

Cyphaspis heissae[123]

Sp. nov

Valid

Van Viersen & Prescher

Middle Devonian

Ahbach Formation

 Germany

An otarionine aulacopleurid, a species of Cyphaspis.

Cyphaspis rommersheimensis[123]

Sp. nov

Valid

Van Viersen & Prescher

Middle Devonian

Freilingen Formation

 Germany

An otarionine aulacopleurid, a species of Cyphaspis.

Cyphaspis walteri[123]

Sp. nov

Valid

Van Viersen & Prescher

Middle Devonian

Bou Dib Formation

 Morocco

An otarionine aulacopleurid, a species of Cyphaspis.

Daketia[124]

Gen. et sp. nov

Valid

Luo et al.

Ordovician (late Tremadocian-earliest Floian)

Tangchi Formation

 China

A member of Asaphida. Genus includes new species D. spinata.

Dalarnepeltis brevifrons[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Tropidocoryphidae.

Diacanthaspis krizi[125]

Sp. nov

Valid

Mergl

Ordovician (late Katian)

Králův Dvůr Formation

 Czech Republic

An odontopleurid, a species of Diacanthaspis.

Dicranogmus pearyi[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Lichidae.

Dionide carlottae[126]

Sp. nov

Valid

Corbacho, Morrison & Ait Addi

Ordovician (Katian)

Ktaoua Formation

 Morocco

A member of Asaphida belonging to the family Dionididae, a species of Dionide.

Dionidella venusta[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Distyrax bibullatus[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Encrinuridae.

Dorypyge gidgealpaensis[114]

Sp. nov

Valid

Sun, Jago & Bentley

Cambrian (Drumian)

Warburton Basin

 Australia

A polymerid trilobite, a species of Dorypyge.

Dulanaspis asper[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Duyunaspis paiwuensis[128]

Sp. nov

Valid

Lei & Peng

Cambrian

Balang Formation

 China

Ectenaspis burkhalteri[115]

Sp. nov

Valid

Amati

Late Ordovician

Welling Formation

 United States

An isoteline asaphid, a species of Ectenaspis.

Ekwanoscutellum agmen[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae.

Elliptocephala jaredi[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Elliptocephala.

Eoharpes yijianfangensis[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Eokosovopeltis expansus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Erratojincella? decora[129]

Sp. nov

Valid

Yuan & Li

Cambrian

Manto Formation

 China

Possibly a species of Erratojincella.

Ganetops[130]

Gen. et comb. nov

Valid

Bignon, Corbacho & López-Soriano

Devonian (late Lochkovian)

 Algeria
 France
 Germany
 Spain

A member of Acastidae belonging to the subfamily Asteropyginae. A new genus for "Asteropyge" ebbae R. and E. Richter (1954); genus also contains "Treveropyge" djemelensis Morzadec (1997), "Treveropyge"? cf. ebbae Smeenk (1983), "Paracryphaeus" gerrinensis Timm (1978) and "Acastella" lata Timm (1978).

Geragnostus (Novoagnostus) contractus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Holmiella domackae[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Holmiella.

Holmiella taurus[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Holmiella.

Hungioides yanjinensis[131]

Sp. nov

Valid

Zhou et al.

Ordovician

 China

A species of Hungioides.

Hystricurus zanderi[132]

Sp. nov

Valid

Adrain et al.

Ordovician (Tremadocian)

 United States

Illaenus cekovioides[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Illaenus tianrongi[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Illaenus usitatus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Iranoleesia (Proasaphiscina?) subconica[129]

Sp. nov

Valid

Yuan & Li

Cambrian

Manto Formation

 China

A species of Iranoleesia.

Isotelus bradleyi[115]

Sp. nov

Valid

Amati

Late Ordovician

Viola Springs Formation

 United States

An isoteline asaphid, a species of Isotelus.

Isotelus skapaneidos[115]

Sp. nov

Valid

Amati

Late Ordovician

Viola Springs Formation
Welling Formation

 United States

An isoteline asaphid, a species of Isotelus.

Isotelus violaensis[115]

Sp. nov

Valid

Amati

Late Ordovician

Viola Springs Formation

 United States

An isoteline asaphid, a species of Isotelus.

Jungosulcus[113]

Gen. et 3 sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae. The type species is J. emilyae; genus also includes new species J. anaphalantos and J. ventricosus.

Kawina sinica[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

A species of Kawina[127] or Cydonocephalus.[133]

Kierarges[134]

Gen. et sp. nov

Valid

Corbacho

Ordovician (Floian)

Fezouata Formation
Santa Justa Formation

 Morocco
 Portugal

A member of Asaphidae. The type species is Kierarges morrisoni.

Lanceaspis[135]

Gen. et sp. nov

Valid

Corbacho

Devonian (Pragian)

The Mader Platform

 Morocco

A member of Acastidae. The type species is Lanceaspis hammondi.

Laneia[113]

Gen. et sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae. The type species is L. enalios.

Ligiscus diana[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae.

Liolalax naresi[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae.

Liostracina tangwangzhaiensis[136]

Sp. nov

Valid

Park et al.

Middle Cambrian

 China

A species of Liostracina.

Lisania changi[114]

Sp. nov

Valid

Sun, Jago & Bentley

Cambrian (Drumian)

Warburton Basin

 Australia

A polymerid trilobite, a species of Lisania.

Lochmanolenellus pentagonalis[137]

Sp. nov

Valid

Mergl

Cambrian (early Dyeran)

North America

A species of Lochmanolenellus.

Lochmanolenellus subquadratus[137]

Sp. nov

Valid

Mergl

Cambrian (early Dyeran)

North America

A species of Lochmanolenellus.

Lochmanolenellus trapezoidalis[137]

Sp. nov

Valid

Mergl

Cambrian (early Dyeran)

North America

A species of Lochmanolenellus.

Lyralichas qiaoi[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Mesonacis wileyi[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Mesonacis.

Minicryphaeus giganteus[130]

Sp. nov

Valid

Bignon, Corbacho & López-Soriano

Devonian (Pragian)

Ihandar Formation

 Morocco

A member of Acastidae belonging to the subfamily Asteropyginae, a species of Minicryphaeus.

Mummaspis delgadoae[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Mummaspis.

Nanillaenus crassus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Nileus amplus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Nileus australis[116]

Sp. nov

Valid

Tortello & Esteban

Early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Nileus.

Olenellus baileyi[119]

Sp. nov

Valid

Gapp & Lieberman

Early Cambrian

Sekwi Formation

 Canada

A member of Olenelloidea, a species of Olenellus.

Olenoides sagittatus[138]

Sp. nov

Valid

Geyer in Geyer et al.

Cambrian (Amgan)

 Kyrgyzstan

A dorypygid corynexochidan, a species of Olenoides.

Opoa limatula[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Scutelluidae.

Owensus[113]

Gen. et sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae. The type species is O. arktoperates.

Parakoldinioidia akerfeldti[139]

Sp. nov

Valid

Westrop & Adrain

Cambrian

Signal Mountain Formation

 United States

A member of Missisquoiidae.

Parakoldinioidia akessoni[139]

Sp. nov

Valid

Westrop & Adrain

Cambrian

Signal Mountain Formation

 United States

A member of Missisquoiidae.

Parakoldinioidia lindgreni[139]

Sp. nov

Valid

Westrop & Adrain

Cambrian

Signal Mountain Formation
Wilberns Formation

 United States

A member of Missisquoiidae.

Parakoldinioidia lopezi[139]

Sp. nov

Valid

Westrop & Adrain

Cambrian

Signal Mountain Formation

 United States

A member of Missisquoiidae.

Parakoldinioidia mendezi[139]

Sp. nov

Valid

Westrop & Adrain

Cambrian

Signal Mountain Formation

 United States

A member of Missisquoiidae.

Paraperaspis[127]

Gen. et 2 sp. et comb. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

A nileid. Genus includes new species P. angustus and P. dawangouensis, as well as "Nileus" convergens Lu (1975).

Paraplethopeltis helli[132]

Sp. nov

Valid

Adrain et al.

Ordovician (Tremadocian)

 United States

Parillaenus exiguus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Parillaenus vetus[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Perryus mikulici[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Encrinuridae.

Profallotaspis tyusserica[140]

Sp. nov

Valid

Bushuev & Goryaeva in Bushuev, Goryaeva & Pereladov

Cambrian

Tyusser Formation

 Russia

A member of Fallotaspididae.

Proetus confluens[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae.

Protolenus (Hupeolenus) bergstroemi[120]

Sp. nov

Valid

Geyer, Buschmann & Elicki

Middle Cambrian

Tröbitz Formation

 Germany

A species of Protolenus.

Pseudinouyia shiliuyuanensis[129]

Sp. nov

Valid

Yuan & Li

Cambrian

Manto Formation

 China

A species of Pseudinouyia.

Pseudoclelandia weymouthae[132]

Sp. nov

Valid

Adrain et al.

Ordovician (Tremadocian)

 United States

Pseudohystricurus wigglesorum[141]

Sp. nov

Valid

Adrain, Karim & Westrop

Ordovician

Garden City Formation

 United States

A dimeropygid trilobite.

Ptychobaba[142]

Gen. et comb. nov

Valid

Pratt & Bordonaro

Cambrian

La Laja Formation
Rome Formation

 Argentina
 United States

A new genus for "Ptychoparella" buttsi Resser (1938).

Punka adamsi[122]

Sp. nov

Valid

McCobb et al.

Early Ordovician

 Greenland

A species of Punka.

Radiurus pauli[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Cheiruridae.

Raragnostus shergoldi[114]

Sp. nov

Valid

Sun, Jago & Bentley

Cambrian (Drumian)

Warburton Basin

 Australia

An agnostoid trilobite, a species of Raragnostus.

Rossaspis leboni[132]

Sp. nov

Valid

Adrain et al.

Ordovician (Tremadocian)

 United States

Sagavia laxa[127]

Sp. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

Sestrostega cylindrica[129]

Sp. nov

Valid

Yuan & Li

Cambrian

Manto Formation

 China

A species of Sestrostega.

Shanxiella (Shanxiella) latilimbata[129]

Sp. nov

Valid

Yuan & Li

Cambrian

Manto Formation

 China

A species of Shanxiella.

Solenoparia gatehousei[114]

Sp. nov

Valid

Sun, Jago & Bentley

Cambrian (Drumian)

Warburton Basin

 Australia

A polymerid trilobite, a species of Solenoparia.

Stegnopsis erythragora[115]

Sp. nov

Valid

Amati

Late Ordovician

Viola Springs Formation
Welling Formation

 United States

An isoteline asaphid, a species of Stegnopsis.

Stegnopsis wellingensis[115]

Sp. nov

Valid

Amati

Late Ordovician

Welling Formation

 United States

An isoteline asaphid, a species of Stegnopsis.

Stenopareia persica[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Illaenidae.

Struveaspis bignoni[143]

Sp. nov

Valid

Corbacho

Devonian (Eifelian)

 Morocco

A member of Phacopidae, a species of Struveaspis.

Tairongia[127]

Gen. et sp. et comb. nov

Valid

Zhou, Yin & Zhou

Ordovician

 China

A lichid. Genus includes new species T. bachuensis, as well as "Lichas" browni Sun (1931).

Trinodus jii[127]

Nom. nov

Zhou, Yin & Zhou

Ordovician

 China

A replacement name for Trinodus latilimbatus Ji (1986).

Winiskia eruga[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae.

Winiskia leptomedia[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae.

Winiskia stickta[113]

Sp. nov

Valid

Hughes & Thomas

Silurian

 Greenland

A member of the family Proetidae.

Other arthropods

Name Novelty Status Authors Age Unit Location Notes Images

Anbarrhacus[144]

Gen. et sp. nov

Valid

Riquelme & Hernández in Riquelme et al.

Early or middle Miocene

 Mexico

A platyrhacid polydesmidan millipede. The type species is Anbarrhacus adamantis.

Annge[145]

Gen. et sp. nov

Valid

Smith et al.

Cambrian (Templetonian)

Giles Creek Dolostone

 Australia

A member of Bradoriida. Genus includes new species A. iperte.

Cambrolongispina[146]

Gen. et 2 sp. nov

Valid

Zhang, Dong & Xiao

Cambrian (Drumian)

 China

A bivalved arthropod of uncertain phylogenetic placement. Genus contains two species: Cambrolongispina reticulata and Cambrolongispina glabra.

Erjiecaris[147]

Gen. et sp. nov

Valid

Fu, Zhang & Budd

Early Cambrian

Chengjiang Lagerstätte

 China

An early bivalved arthropod of uncertain phylogenetic placement. The type species is Erjiecaris minusculo.

Falcatamacaris[148]

Gen. et sp. nov

Valid[149]

Ortega-Hernández et al.

Cambrian (Guzhangian)

Weeks Formation

 United States

An artiopodan arthropod. The type species is Falcatamacaris bellua.

Flemingopsis anteriospinata[145]

Sp. nov

Valid

Smith et al.

Cambrian (Templetonian)

Giles Creek Dolostone

 Australia

A member of Bradoriida.

Haifengella[150]

Gen. et sp. nov

Valid

Zhao et al.

Early Cambrian

Chengjiang Lagerstätte

 China

An arthropod related to Helmetia. The type species is Haifengella corona.

Jiucunella phaseloa[151]

Sp. nov

Valid

Betts et al.

Early Cambrian

 Australia

A member of Bradoriida, a species of Jiucunella.

Jixinlingella daimonikoa[151]

Sp. nov

Valid

Betts et al.

Early Cambrian

 Australia

A member of Bradoriida, a species of Jixinlingella.

Limulus darwini[152]

Sp. nov

Valid

Kin & Błażejowski

Late Jurassic (late Tithonian)

Kcynia Formation

 Poland

A horseshoe crab. Originally described as a species of Limulus; Bicknell et al. (2021) transferred this species to the genus Crenatolimulus.[153]

Maatidesmus[144]

Gen. et sp. nov

Valid

Riquelme & Hernández in Riquelme et al.

Early or middle Miocene (c. 23–15 Mya)

Mexican amber

 Mexico

A chelodesmid polydesmidan millipede. The type species is M. paachtun.

Monasterium ferox[145]

Sp. nov

Valid

Smith et al.

Cambrian (Templetonian)

Giles Creek Dolostone

 Australia

A member of Bradoriida.

Mongolitubulus anthelios[151]

Sp. nov

Valid

Betts et al.

Early Cambrian

 Australia

A member of Bradoriida, a species of Mongolitubulus.

Mongolitubulus tunpere[145]

Sp. nov

Valid

Smith et al.

Cambrian (Templetonian)

Giles Creek Dolostone

 Australia

A member of Bradoriida.

Neokunmingella moroensis[151]

Sp. nov

Valid

Betts et al.

Early Cambrian

 Australia

A member of Bradoriida, a species of Neokunmingella.

Polzia[154]

Gen. et sp. nov

Valid

Hegna, Vega & González-Rodríguez

Cretaceous (Albian to Cenomanian)

El Doctor Formation

 Mexico

A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans). The type species is Polzia eldoctorensis.

Sinskolutella cuspidata[151]

Sp. nov

Valid

Betts et al.

Early Cambrian

 Australia

A member of Bradoriida, a species of Sinskolutella.

Thylacares[155]

Gen. et sp. nov

Valid

Haug et al.

Silurian (late Telychian)

Brandon Bridge Formation

 United States

A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans). The type species is Thylacares brandonensis.

Victoriacaris[154]

Gen. et sp. nov

Valid

Hegna, Vega & González-Rodríguez

Cretaceous (Albian to Cenomanian)

El Doctor Formation

 Mexico

A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans. The type species is Victoriacaris muhiensis.

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