2014 in arthropod paleontology

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This list of fossil arthropods described in 2014 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2014. The list only includes taxa at the level of genus or species.

Arachnids

Newly named taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Baalzebub? mesozoicum[1]	Sp. nov	Valid	Penney	Cretaceous (middle Cenomanian to early Santonian)		France	A spider of uncertain placement. Originally described as a ray spider; Wunderlich & Müller (2020) transferred it to the family Zarqaraneidae.[2]
Cheirolepidoptus[3]	Gen. et sp. nov	Valid [4]	Sidorchuk & Lindquist in Sidorchuk et al.	Late Triassic (Carnian)		Italy	A Tetrapodili mite The type species is C. dolomiticus.
Feaella (Tetrafeaella) groehni[5]	Sp. nov	Valid	Henderickx in Henderickx & Boone	Eocene	Baltic amber	Europe	A feaellid pseudoscorpion
Hastocularis[6]	Gen. et sp. nov	Valid	Garwood et al.	Carboniferous (late Stephanian)		France	A Tetrophthalmi Opiliones. The type species is H. argus.
Kronocharon[7]	Gen. et sp. nov	Valid	Engel & Grimaldi	Cretaceous (latest Albian or earliest Cenomanian)	Burmese amber	Myanmar	An Amblypygi originally assigned to Neoamblypygi excluded from this group by Garwood et al., 2017[8]). The type species is Kronocharon prendinii.
Mayachernes[9]	Gen. et sp. nov	Jr synonym	Riquelme et al.	Miocene	Mexican amber	Mexico	A chernetid pseudoscorpion. The type species is Mayachernes maatiatus. Moved to Byrsochernes maatiatus by Judson (2016)[10]
Minyacarus[3]	Gen. et sp. nov	Valid[4]	Sidorchuk & Lindquist in Sidorchuk et al.	Late Triassic (Carnian)		Italy	A Tetrapodili mite. The type species is M. aderces.
Montsecarachne[11]	Gen. et sp. nov	Valid	Selden	Early Cretaceous (c. 129 Ma)	El Montsec	Spain	A plectreurid spider. The type species is M. amicus.
?Opilioacarus groehni[12]	Sp. nov	Valid	Dunlop & Bernardi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An opilioacarid mite.
Paracharonopsis[7]	Gen. et sp. nov	Valid	Engel & Grimaldi	Eocene (Ypresian)	Cambay Shale Formation	India	A Paracharontidae Amblypygi. The type species is P. cambayensis.
Paratrombium rovniense[13]	Sp. nov	Valid	Konikiewicz & Mąkol	Eocene	Rovno amber	Ukraine	A mite.
Pleophrynus hawesi[14]	Sp. nov	Valid	Dunlop et al.	Carboniferous (Pennsylvanian)	Astrasado Formation	United States	An eophrynid trigonotarbid.
Pulchellaranea[15]	Gen. et sp. nov	Valid	Poinar	Cenozoic	Dominican amber	Dominican Republic	An orb-weaver spider. The type species is P. pedunculata.
Seppo[16]	Gen. et sp. nov	Valid	Selden & Dunlop	Early Jurassic (Toarcian)	Ciechocinek Formation	Germany	A probable palpimanoidean spider. The type species is S. koponeni.
Tityus knodeli[17]	Sp. nov	Valid	Lourenço	Miocene	Mexican amber	Mexico	A buthid scorpion.
Tuckerella fossilibus[18]	Sp. nov	Valid	Khaustov, Sergeyenko & Perkovsky	Eocene	Rovno amber	Ukraine	A peacock mite.

Newly named crustaceans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Acanthaxius carinatus[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidae, a species of Acanthaxius.
Acanthochirana liburiaensis[20]	Sp. nov	Valid	Garassino, Audo, Charbonnier & Schweigert in Bravi et al.	Middle Jurassic (Bajocian/Bathonian)		Italy	A member of Aegeridae, a species of Acanthochirana.
Acanthochirana triassica[21]	Sp. nov	Valid	Garassino, Schweigert & Muscio	Late Triassic (Norian)	Dolomia di Forni Formation	Italy	A member of Aegeridae, a species of Acanthochirana.
Adamanteryon[22]	Gen. et sp. nov	Valid	Audo et al.	Middle Jurassic	La Voulte-sur-Rhône Lagerstätte	France	A member of Polychelida. The type species is Adamanteryon fourneti.
Alavatanais margulisae[23]	Sp. nov	Valid[24]	Sánchez-García, Peñalver & Delclòs in Sánchez-García et al.	Early Cretaceous		Spain	A paratanaoid tanaid, a species of Alavatanais.
Alessandranina[25]	Gen. et comb. nov	Valid	Karasawa et al.	Late Cretaceous (Cenomanian)		United Kingdom	A palaeocorystid crab. The type species is "Notopocorystes" (Cretacoranina) ornatus Wright and Collins (1972).
Ammopylocheles petersi[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Ammopylocheles.
Anisaeger[27]	Gen. et 2 sp. nov	Valid	Schweitzer et al.	Middle Triassic (Anisian)		China	A member of Aegeridae. The type species is Anisaeger brevirostrus; genus also contains Anisaeger spiniferus.
Annuntidiogenes jurassicus[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Annuntidiogenes.
Antrimpos colettoi[21]	Sp. nov	Valid	Garassino, Schweigert & Muscio	Late Triassic (Norian)	Dolomia di Forni Formation	Italy	A member of Penaeidae. Originally described as a species of Antrimpos, but subsequently transferred to the genus Ladinicaris.[28]
Araripenaeus[29]	Gen. et sp. nov	Valid	Pinheiro, Saraiva & Santana	Early Cretaceous (Albian)	Romualdo Formation	Brazil	A member of Penaeoidea of uncertain phylogenetic placement. The type species is Araripenaeus timidus.
Arcania kobayashinobuakii[30]	Sp. nov	Valid	Karasawa	Middle Pleistocene	Toyohashi Formation	Japan	A member of Leucosiidae, a species of Arcania.
Archaeoniscus italiensis[31]	Sp. nov	Valid	Jones, Feldmann & Garassino	Jurassic		Italy	An isopod, a species of Archaeoniscus.
Ariecornibus[32]	Gen. et sp. nov	Valid	Franţescu	Early Cretaceous (late Albian)		United States	A crab. The type species is Ariecornibus schweitzerae.
Axiopsis pawpawensis[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidae, a species of Axiopsis.
Axiopsis sampsonumae[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidae, a species of Axiopsis.
Axiopsis spinifera[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidae, a species of Axiopsis.
Bellcarcinus[33]	Gen. et sp. nov	Valid	Luque	Early Cretaceous (late Aptian)	Paja Formation	Colombia	A crab belonging to the group Raninoidia, possibly a member of the family Orithopsidae. The type species is Bellcarcinus aptiensis.
Belos[32]	Gen. et sp. nov	Valid	Franţescu	Early Cretaceous (late Albian)		United States	A crab. The type species is Belos trispinae.
Bolbozoe beccata[34]	Sp. nov	Valid[35]	Perrier et al.	Early Silurian		Australia	A myodocope ostracod, a species of Bolbozoe.
Branchiocarcinus pacificus[36]	Sp. nov	Valid	Nyborg, Ossó & Vega	Late Cretaceous (Maastrichtian)	Moreno Formation	United States	A portunoid crab. Originally described as a member of the family Icriocarcinidae and a species of Branchiocarcinus; Kornecki, Feldmann & Schweitzer (2017) considered the family Icriocarcinidae to be a junior synonym of the family Lithophylacidae and transferred B. pacificus to the genus Lithophylax.[37]
Brazilestheria[38]	Gen. et comb. nov	Valid	Chen & Shen	Early Cretaceous		Brazil	A clam shrimp belonging to the family Sinoestheriidae. Genus includes "Asmussia" souzae Guerin-Franiatte & Taquet (1993).
Canalarta[34]	Gen. et sp. nov	Valid[35]	Perrier et al.	Early Silurian		Australia	A myodocope ostracod. The type species is Canalarta papata.
Candonopsis alagoensis[39]	Sp. nov	Valid	Tomé, Lima Filho & Neumann	Early Cretaceous	Araripe basin	Brazil	An ostracod.
Cantabroxanthus[40]	Gen. et sp. nov	Valid	Ossó & Díaz Isa	Late Cretaceous (middle Campanian)		Spain	A feldmanniid crab, a relative of etyiids. The type species is Cantabroxanthus loredoensis.
Casertanus[20]	Gen. et sp. nov	Valid	Garassino, Audo, Charbonnier & Schweigert in Bravi et al.	Middle Jurassic (Bajocian/Bathonian)		Italy	A member of Sergestidae. The type species is Casertanus sabellicus.
Ceratobairdia crassispinosa[41]	Sp. nov	Valid	Monostori & Tóth	Triassic		Hungary	An ostracod, a species of Ceratobairdia.
Claudioranina[25]	Gen. et comb. nov	Valid	Karasawa et al.	Eocene (Lutetian-Bartonian)		Italy	A raninid crab. The type species is "Cyrtorhina" oblonga Beschin, Busulini, De Angeli & Tessier (1988).
Clinocypris epacrus[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Clinocypris.
Colombicarcinus[25]	Gen. et sp. nov	Valid	Karasawa et al.	Early Cretaceous (Aptian)		Colombia	A necrocarcinid crab. The type species is Colombicarcinus laevis.
Concinnalepas[43]	Gen. et comb. nov	Valid	Gale	Late Jurassic		United Kingdom	A zeugmatolepadid barnacle. A new genus for "Pollicipes" concinnus Morris (1845).
Cophinia grekoffi[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Cophinia.
Cophinia ovalis[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Cophinia.
Corallomursia[46]	Gen. et sp. nov	Valid	De Angeli & Ceccon	Early Eocene		Italy	A crab belonging to the family Calappidae. Genus includes new species C. eocaena.
Ctenocheles fritschi[47]	Sp. nov	Valid	Hýžný, Kočová Veselská & Dvořák	Late Cretaceous (Coniacian)	Březno Formation	Czech Republic	A ctenochelid axiidean, a species of Ctenocheles.
Cycleryon romani[22]	Sp. nov	Valid	Audo et al.	Middle Jurassic	La Voulte-sur-Rhône Lagerstätte	France	A member of Polychelida, a species of Cycleryon.
Cymothoidana[48]	Gen. et sp. nov	Valid	Jarzembowski et al.	Early Cretaceous	Wealden Supergroup	United Kingdom	A cymothoidan isopod. The type species is Cymothoidana websteri.
Cypria poietes[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Cypria.
Cytheropteron paramassoni[49]	Sp. nov	Valid	Yasuhara & Okahashi	Late Quaternary		North Atlantic Ocean (Rockall Plateau)	A cytherurid podocopid ostracod, a species of Cytheropteron.
Damonella grandiensis[39]	Sp. nov	Valid	Tomé, Lima Filho & Neumann	Early Cretaceous	Araripe basin	Brazil	An ostracod.
Dawsonius tigris[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidea belonging to the family Ctenochelidae, a species of Dawsonius.
Devonostenopus[50]	Gen. et sp. nov	Valid	Jones et al.	Devonian	Huntley Mountain Formation	United States	A member of Stenopodidea. The type species is Devonostenopus pennsylvaniensis.
Dicerobairdia latispinosa[41]	Sp. nov	Valid	Monostori & Tóth	Triassic		Hungary	An ostracod, a species of Dicerobairdia.
Diogenicheles[51]	gen et sp nov	Valid	Fraaije et al.	Oxfordian		Poland	A parapylochelid hermit crab. The type species is Diogenicheles theodorae.
Distaeger[27]	Gen. et sp. nov	Valid	Schweitzer et al.	Middle Triassic (Anisian)		China	A member of Aegeridae. The type species is Distaeger prodigiosus.
Distefania(?) tangishirazensis[52]	Sp. nov	Valid	Jagt et al.	Early Cretaceous (Albian)	Kazhdumi Formation	Iran	A member of Dromioidea, possibly a species of Distefania.
Distefania(?) vanrijsselti[53]	Sp. nov	Valid	Jagt, Fraaije & van Bakel	Late Cretaceous (late Maastrichtian)	Maastricht Formation	Netherlands	A member of Dromioidea, possibly a species of Distefania.
Dromia evae[54]	Sp. nov	Valid	Collins	Miocene		Austria	A member of Dromiidae, a species of Dromia.
Dromiopsis paleogenicus[46]	Sp. nov	Valid	De Angeli & Ceccon	Early Eocene		Italy	A crab belonging to the family Dynomenidae.
Dundgobiestheria[55]	Gen. et sp. nov	Valid	Li in Li et al.	Probably Middle Jurassic	Eedemt Formation	Mongolia	A clam shrimp. The type species is Dundgobiestheria mandalgobiensis.
Electrotanais[23]	Gen. et sp. nov	Valid[24]	Sánchez-García, Peñalver & Delclòs in Sánchez-García et al.	Early Cretaceous		Spain	A paratanaoid tanaid. The type species is Electrotanais monolithus.
Entomozoe bispinosa[56]	Sp. nov	Valid	Wang & Wu	Early Silurian	Shimenkan Formation	China	An ostracod, a species of Entomozoe.
Entomozoe suboblonga[56]	Sp. nov	Valid	Wang & Wu	Early Silurian	Shimenkan Formation	China	An ostracod, a species of Entomozoe.
Entomozoe wukeensis[56]	Sp. nov	Valid	Wang & Wu	Early Silurian	Shimenkan Formation	China	An ostracod, a species of Entomozoe.
Eogordonella[57]	Gen. et sp. nov	Valid	Garassino et al.	Eocene		Iran	A solenocerid penaeoid. The type species is Eogordonella iranianiensis.
Eomaldivia? friebei[54]	Sp. nov	Valid	Collins	Miocene		Austria	A member of Trapeziidae, possibly a species of Eomaldivia.
Eonatatolana[58]	Gen. et sp. nov	Valid[59]	Etter	Middle Jurassic		Germany	A cirolanid isopod. The type species is Eonatatolana geisingensis.
Eopabdehus[57]	Gen. et sp. nov	Valid	Garassino et al.	Eocene		Iran	A penaeid. The type species is Eopabdehus babaheydariensis.
Eosestheria lushangfenensis[60]	Sp. nov	Valid	Liao, Shen & Huang	Early Cretaceous	Lushangfen Formation	China	A clam shrimp, a species of Eosestheria.
Epibrachylepas[61]	Gen. et sp. et comb. nov	Valid[62]	Gale in Gale & Sørensen	Late Cretaceous (Campanian and Maastrichtian)		Netherlands Sweden	A basal balanomorph barnacle. The type species is Epibrachylepas newmani; genus also includes Epibrachylepas smeetsi (Bosquet, 1857).
Eryma osciensis[20]	Sp. nov	Valid	Garassino, Audo, Charbonnier & Schweigert in Bravi et al.	Middle Jurassic (Bajocian/Bathonian)		Italy	A member of the family Erymidae.
Etcheslepas[43]	Gen. et sp. et comb. nov	Valid	Gale	Late Jurassic		United Kingdom	A member of Pollicipedidae. Genus contains the new species Etcheslepas durotrigensis, as well as "Zeugmatolepas" fragilis Withers (1928).
Etisus evamuellerae[63]	Sp. nov	Valid	Hyžný, van Bakel & Guinot	Middle Miocene		Austria Hungary Spain?	A member of Xanthidae, a species of Etisus.
Eucytherura myrsinae[64]	Sp. nov	Valid	Uffenorde in Uffenorde & Radtke	Oligocene		Germany	A cytherurid podocopid ostracod.
Eucytherura zehali[49]	Nom. nov	Valid	Yasuhara & Okahashi	Late Quaternary		North Atlantic Ocean (Carolina Slope)	A cytherurid podocopid ostracod; a replacement name for Eucytherura hazeli Yasuhara, Okahashi & Cronin (2009).
Eukloedenella adcapitisdolorella[65]	Sp. nov	Valid[66]	Forel	Early Triassic	Kokarkuyu Formation	Turkey	An ostracod, a species of Eukloedenella.
Eurotanais[23]	Gen. et sp. nov	Valid[24]	Sánchez-García, Peñalver & Delclòs in Sánchez-García et al.	Early Cretaceous		Spain	A paratanaoid tanaid. The type species is Eurotanais terminator.
Falklandicypris[42]	Gen. et 2 sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod. The type species is Falklandicypris petrasaltata; genus also contains Falklandicypris desiderata.
Faxelepas[67]	Gen. et comb. nov	Valid[68]	Gale	Late Cretaceous (Maastrichtian) to late Miocene[69]		Denmark Italy Netherlands Spain[69] Sweden	A barnacle related to verrucomorphs. A new genus for "Pycnolepas" bruennichi Withers (1914); genus also includes "Pycnolepas" paronai De Alessandri (1895).[69]
Fossocytheridea potiguarensis[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Fossocytheridea.
Fossocytheridea tiberti[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Fossocytheridea.
Gangamoncythere[42]	Gen. et sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod. The type species is Gangamoncythere colini.
Glabropilumnus nitidus[54]	Sp. nov	Valid	Collins	Miocene		Austria	A member of Pilumnidae, a species of Glabropilumnus.
Globotalicypridea mirabilis[70]	Sp. nov	Valid	Silye, Colin & Codrea	Late Cretaceous (Maastrichtian)		Romania	A talicyprideine cypridid ostracod, a species of Globotalicypridea.
Gracilimanus[71]	Gen. et sp. nov	Valid	Feldmann & Schweitzer in Guyer, Schweitzer & Feldmann	Middle Jurassic (Bajocian/Bathonian)	Gypsum　Springs Formation	United States	A member of Astacidea, possibly a member of the family Stenochiridae. The type species is Gracilimanus obscurus.
Giulianolyreidus[25]	Gen. et comb. nov	Valid	Karasawa et al.	Paleocene (Thanetian)		United States	A lyreidid crab. The type species is "Symethis" johnsoni Rathbun (1935); genus also contains "Symnista" bidentata Rathbun (1935).
Gypsonicus[71]	Gen. et sp. nov	Valid	Schweitzer & Feldmann in Guyer, Schweitzer & Feldmann	Middle Jurassic (Bajocian/Bathonian)	Gypsum　Springs Formation	United States	A member of Astacidea belonging to the family Stenochiridae. The type species is Gypsonicus wyomingensis.
Harpactocarcinus fedrigoi[72]	Sp. nov	Valid	Beschin, De Angeli & Zorzin	Eocene		Italy	A zanthopsid carpilioid crab, a species of Harpactocarcinus.
Haughtonileberis dinglei[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian to Santonian)		Brazil Nigeria	A trachyleberidid ostracod, a species of Haughtonileberis.
Hemicytherura viviersae[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherurid ostracod, a species of Hemicytherura.
Hepatus pauli[73]	Sp. nov	Valid	Collins, Garvie & Mellish	Pleistocene	Beaumont Formation	United States	A member of Aethridae, a species of Hepatus.
Hourcqia woodi[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Hourcqia.
Housacheles ovalis[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Housacheles.
Hyas chippubetsuensis[74]	Sp. nov	Valid	Kato, Nakashima & Yanagisawa	Early Pliocene	Chippubetsu Formation	Japan	A member of Oregoniidae, a species of Hyas.
Hyas tentokujiensis[74]	Sp. nov	Valid	Kato, Nakashima & Yanagisawa	Early Pliocene	Tentokuji Formation	Japan	A member of Oregoniidae, a species of Hyas.
Icenilepas[43]	Gen. et sp. nov	Valid	Gale	Late Cretaceous (Santonian)		United Kingdom	A zeugmatolepadid barnacle. The type species is Icenilepas collinsi.
Ilhasina? leiodermatus[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, possibly a species of Ilhasina.
Ilyocypris bisulcata[75]	Sp. nov	Valid	Qu et al.	Early Paleocene	Mingshui Formation	China	An ilyocypridid cypridoid ostracod, a species of Ilyocypris.
Italianira[25]	Gen. et comb. nov	Valid	Karasawa et al.	Eocene (Ypresian)		Italy	A raninid crab. The type species is "Ranilia" punctulata Beschin, Busulini, De Angeli & Tessier (1988).
Jandairella[44]	Gen. et sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A trachyleberidid ostracod. The type species is Jandairella obesa.
Kegelina[76]	Gen. et comb. nov	Valid	de Queiroz Neto, Sames & Colin	Early Cretaceous		Brazil Gabon United States	A cypridoid ostracod. A new genus for "Cypridea" anomala Peck (1941), "Cypridea" armata Krömmelbein (1962), "Cypridea" bisculpturata Wicher (1959), "Cypridea" depressa Moura (1972) and "Cypridea" kegeli Wicher (1959).
Kotoracythere paiki[77]	Sp. nov	Valid	Lee	Pliocene-Pleistocene	Seogwipo Formation	South Korea	An ostracod, a species of Kotoracythere.
Krausella danukalovae[78]	Sp. nov	Valid	Melnikova & Danukalova	Ordovician		Russia	An ostracod, a species of Krausella.
Kuzminaella arabieni[79]	Sp. nov	Valid	Groos-Uffenorde in Groos-Uffenorde & Rabien	Devonian		Germany	An ostracod.
Lepidophthalmus paratethyensis[80]	Sp. nov	Valid[81]	Gašparič & Hyžný	Early Miocene (late Burdigalian)		Slovenia	A member of Callianassidae, a species of Lepidophthalmus.
Longiscula kazarkina[78]	Sp. nov	Valid	Melnikova & Danukalova	Ordovician		Russia	An ostracod, a species of Longiscula.
Looneyellopsis tuberculatus[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Looneyellopsis.
Loxocorniculum? narendrai[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A loxoconchid ostracod, possibly a species of Loxocorniculum.
Luprisca[82]	Gen. et sp. nov	Valid	Siveter et al.	Ordovician (Katian)	Lorraine Group	United States	A myodocope ostracod. The type species is Luprisca incuba.
Martillepas[43]	Gen. et comb. nov	Valid	Gale	Late Jurassic		United Kingdom	A zeugmatolepadid barnacle. A new genus for "Calantica" ovalis Withers (1928), "Zeugmatolepas" costata Withers (1928) and "Zeugmatolepas" hollisi Withers (1928).
Masticacheles[51]	Gen et 2 sp nov	Valid	Fraaije et al.	Oxfordian to Kimmeridgian		Germany Poland	A parapylochelid hermit crab. The type species is Masticacheles longirostris from Oxfordian of Poland;[51] subsequently a second species Masticacheles minimus from late Kimmeridgian of Germany was named by Fraaije (2014).[26]
Mesoparapylocheles jaegeri[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Mesoparapylocheles.
Mesoparapylocheles schweigerti[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Mesoparapylocheles.
Monoceratina hussonae[65]	Sp. nov	Valid[66]	Forel	Early Triassic	Kokarkuyu Formation	Turkey	An ostracod, a species of Monoceratina.
Montemagrellus[46]	Gen. et sp. nov	Valid	De Angeli & Ceccon	Early Eocene		Italy	A crab belonging to the family Crossotonotidae. Genus includes new species M. denticulatus.
Muelleristhes[83]	Gen. et comb. nov	Valid	Garassino, De Angeli & Pasini	Late Cretaceous (Cenomanian-Turonian)		Morocco	A porcelain crab; a new genus for "Paragalathea" africana Garassino, De Angeli & Pasini (2008).
Munidopsis palmuelleri[84]	Sp. nov	Valid	Hyžný et al.	Middle Miocene	Slovenske Gorice Formation	Slovenia	A munidopsid galatheoid, a species of Munidopsis.
Musacchiocythere[42]	Gen. et comb. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod. The type species is Musacchiocythere sarunata (Musacchio, 1978).
Neopilumnoplax pohorjensis[80]	Sp. nov	Valid[81]	Gašparič & Hyžný	Early Miocene (late Burdigalian)		Slovenia	A member of Mathildellidae, a species of Neopilumnoplax.
Nothocarapacestheria[85]	Gen. et sp. nov	Valid	Rohn, Dutra & Cabral	Probably Jurassic	Caturrita Formation	Brazil	A clam shrimp belonging to the family Eosestheriidae. The type species is Nothocarapacestheria soturnensis.
Occultocaris[86]	Gen. et sp. nov	Valid	Winkler	Late Jurassic (early Tithonian)	Solnhofen Plattenkalk	Germany	A member of Caridea, possibly a bresilioid. The type species is Occultocaris frattigianii.
Ovocytheridea anterocompressa[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Ovocytheridea.
Ovocytheridea posteroprojecta[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Ovocytheridea.
Ovocytheridea reymenti[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous		Brazil Nigeria	A cytherideid ostracod, a species of Ovocytheridea.
Ovocytheridea triangularis[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Ovocytheridea.
Pagurus retznensis[54]	Sp. nov	Valid	Collins	Miocene		Austria	A hermit crab, a species of Pagurus.
Pagurus texensis[32]	Sp. nov	Valid	Franţescu	Early Cretaceous (late Albian)		United States	A hermit crab, a species of Pagurus.
Palaega furcillanatis[31]	Sp. nov	Valid	Jones, Feldmann & Garassino	Jurassic		Italy	An isopod, a species of Palaega.
Palaeogammarus debroyeri[87]	Sp. nov	Valid	Jażdżewski, Grabowski & Kupryjanowicz	Eocene		Probably Russia (Kaliningrad Oblast)	A member of Amphipoda found in Baltic amber, a species of Palaeogammarus.
Palinurellus bericus[88]	Sp. nov	Valid	De Angeli & Garassino	Eocene (Priabonian)		Italy	A furry lobster, a species of Palinurellus.
Palladiocarcinus[46]	Gen. et sp. nov	Valid	De Angeli & Ceccon	Early Eocene		Italy	A crab belonging to the family Pilumnidae. Genus includes new species P. brevidentatus.
Palmunidopsis[89]	Gen. et sp. nov	Valid	Fraaije	Miocene (Tortonian)	Pakhna Formation	Cyprus	A munidopsid galatheoid. The type species is Palmunidopsis muelleri.
Parabrachylepas[61]	Gen. et comb. nov	Valid[62]	Gale in Gale & Sørensen	Late Cretaceous (Campanian)		Sweden	A basal balanomorph barnacle. The type species is Parabrachylepas ifoensis (Withers, 1935).
Paracypris ? krivipotokensis[90]	Sp. nov	Valid	Forel in Sudar et al.	Early Triassic (Olenekian)	Obnica Formation	Serbia	A cypridoid ostracod belonging to the family Paracyprididae, possibly a species of Paracypris.
Paracypris vietnamensis[91]	Sp nov	Valid	Tanaka & Komatsu in Shigeta et al.	Early Triassic (Olenekian)	Bac Thuy Formation	Vietnam	An ostracod, a species of Paracypris.
Paraplesiocypridea[42]	Gen. et sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod. The type species is Paraplesiocypridea alloios.
Parapylochelitergites deductius[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Parapylochelitergites.
Parapylochelitergites prepustulosus[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Parapylochelitergites.
Paraxiopsis erugatus[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidae, a species of Paraxiopsis.
Parsacus[57]	Gen. et sp. et comb. nov	Valid	Garassino et al.	Eocene		Iran Italy	A slipper lobster. Genus contains new species Parsacus eocenicus, as well as "Parribacus" cristatus Förster (1984).
Pauline nivisis[92]	Sp. nov	Valid	Perrier et al.	Silurian (late Telychian)	Pentamerus Bjerge Formation	Greenland	A cylindroleberidid myodocope ostracod, a species of Pauline.
Paulinecaris[93]	Gen. et sp. nov	Valid[94]	Waloszek et al.	Cambrian		Sweden	An entomostracan of uncertain phylogenetic placement. The type species is Paulinecaris siveterae.
Pedupycnolepas[67]	Gen. et comb. nov	Valid[68]	Gale	Early Cretaceous (probably early Aptian)		Antarctica (Alexander Island)	A barnacle; a new genus for "Pycnolepas" articulata Collins (1980).
Periacanthus tetracornis[95]	Sp. nov	Valid	Ferratges et al.	Eocene		Spain	A crab belonging to the family Epialtidae.
Perissocytheridea caudata[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Perissocytheridea.
Perissocytheridea jandairensis[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Perissocytheridea.
Perissocytheridea mossoroensis[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherideid ostracod, a species of Perissocytheridea.
Petrochirus sanctilazzari[96]	Sp. nov	Valid	Baldanza et al.	Early Pleistocene		Italy	A diogenid hermit crab, a species of Petrochirus.
Petrolisthes albianicus[32]	Sp. nov	Valid	Franţescu	Early Cretaceous (late Albian)		United States	Originally classified as a porcelain crab belonging to the genus Petrolisthes. Robins & Klompmaker (2019) transferred it to the genus Hispanigalathea belonging to the galatheoid family Catillogalatheidae.[97]
Pilgrimcheles[51]	gen et 3 sp nov	Valid	Fraaije et al.	Oxfordian to Kimmeridgian		Germany Poland	A parapylochelid hermit crab. The type species is Pilgrimcheles karolinae from Oxfordian of Poland;[51] subsequently additional two species, Pilgrimcheles kersteni and Pilgrimcheles vonmeyeri, were described from Kimmeridgian of Germany by Fraaije (2014).[26]
Pisidia? subnodosa[54]	Sp. nov	Valid	Collins	Miocene		Austria	A porcelain crab, possibly a species of Pisidia.
Planobranchia palmuelleri[98]	Sp. nov	Valid	Artal, van Bakel & Onetti	Eocene (Lutetian)	Coll de Malla Formation	Spain	A member of Inachidae, a species of Planobranchia.
Plioaxius texensis[19]	Sp. nov	Valid	Frantescu	Early Cretaceous (Albian)	Paw Paw Formation	United States	A member of Axiidae, a species of Plioaxius.
Portunus muelleri[54]	Sp. nov	Valid	Collins	Miocene		Austria	A member of Portunidae, a species of Portunus.
Potiguarella[44]	Gen. et 2 sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)		Brazil Gabon	A trachyleberidid ostracod. The type species is Potiguarella grosdidieri; genus also contains Potiguarella coimbrai.
Priscoverruca[67]	Gen. et comb. et sp. nov	Valid[68]	Gale	Late Cretaceous		Belgium Germany Netherlands United Kingdom	A barnacle. The type species is "Verruca" prisca Bosquet (1854); genus also includes new species P. elongata.
Procytherura ballentae[44]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Turonian)	Potiguar Basin	Brazil	A cytherurid ostracod, a species of Procytherura.
Protocosta babinoti[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A trachyleberidid ostracod, a species of Protocosta.
Proverruca dentifer[67]	Sp. nov	Valid[68]	Gale	Late Cretaceous (Campanian)		Belgium Sweden	A barnacle.
Pylochelitergites exspectatus[99]	Sp. nov	Valid	Fraaije et al.	Late Jurassic (Oxfordian)		Poland	A member of Pylochelidae, a species of Pylochelitergites.
Pyreneplax[100]	Gen. et sp. et comb. nov	Valid	Ossó, Domínguez & Artal	Eocene		Italy Spain United States	A member of Vultocinidae. The type species is P. basaensis; genus also includes P. ranosa (Beschin, Busulini, De Angeli & Tessier, 2002), P. sandersi (Blow & Manning, 1997) and P. sommarugai (Beschin, Busulini & Tessier, 2009).
Quadratoplanus[32]	Gen. et sp. nov	Valid	Franţescu	Early Cretaceous (late Albian)		United States	A crab. The type species is Quadratoplanus primitivus.
Retropluma slovenica[80]	Sp. nov	Valid[81]	Gašparič & Hyžný	Early Miocene (late Burdigalian)		Slovenia	A member of Retroplumidae, a species of Retropluma.
Reviya sylvieae[65]	Sp. nov	Valid[66]	Forel	Early Triassic	Kokarkuyu Formation	Turkey	An ostracod, a species of Reviya.
Schramidontus[101]	Gen. et sp. nov	Valid	Gueriau, Charbonnier & Clément	Late Devonian (Famennian)		Belgium	A member of Eucarida related to Angustidontus. The type species is Schramidontus labasensis.
Scututergites[26]	Gen. et sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab. The type species is Scututergites anteroindentatus.
Semicytherura musacchioi[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A cytherurid ostracod, a species of Semicytherura.
Sphaeroma montefallanoense[31]	Sp. nov	Valid	Jones, Feldmann & Garassino	Jurassic		Italy	An isopod, a species of Sphaeroma.
Stagmacaris subcircularis[26]	Sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab, a species of Stagmacaris.
Stevea martini[102]	Sp. nov	Valid	Feldmann, Schweitzer & Portell	Early Paleocene	Clayton Formation	United States	A member of Hexapodidae, a species of Stevea.
Synurella aliciae[87]	Sp. nov	Valid	Jażdżewski, Grabowski & Kupryjanowicz	Eocene		Probably Russia (Kaliningrad Oblast)	A member of Amphipoda found in Baltic amber, a species of Synurella.
Tamnacythere[77]	Gen. et sp. nov	Valid	Lee	Pliocene-Pleistocene	Seogwipo Formation	South Korea	An ostracod. The type species is Tamnacythere seogwipoensis.
Tanycythere[103]	Gen. et 6 sp. nov	Valid	Cabral et al.	Early Jurassic to Early Cretaceous (Hettangian to Barremian?)		Western and central Europe	An ostracod. Genus contains six species: T. caudata, T. praecaudata (with two subspecies: T. praecaudata praecaudata and T. praecaudata parallela), T. duartei, T. posteroelongata, T. wattonensis and T. procera.
Tealliocaris walloniensis[104]	Sp. nov	Valid[105]	Gueriau, Charbonnier & Clément	Late Devonian (Famennian)		Belgium	A pygocephalomorph crustacean,[106] a species of Tealliocaris.
Telamonocarcinus antiquus[107]	Sp. nov	Valid[108]	Luque	Early Cretaceous (early Albian)		Colombia	A crab, a species of Telamonocarcinus.
Tesseropora canariana[109]	Sp. nov	Valid	Hornung	Pliocene (Zanclean)		Spain ( Canary Islands)	A tetraclitid balanomorph barnacle, a species of Tesseropora.
Tethyseryon[20]	Gen. et sp. nov	Valid	Garassino, Audo, Charbonnier & Schweigert in Bravi et al.	Middle Jurassic (Bajocian/Bathonian)		Italy	A member of Eryonidae. The type species is Tethyseryon campanicus.
Theriosynoecum ballentae[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Theriosynoecum.
Theriosynoecum petasmathylacus[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Theriosynoecum.
Tianzhuestheria jianchangensis[110]	Sp. nov	Valid	Wang	Middle–Late Jurassic		China	A clam shrimp.
Timiriasevia fluitans[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Timiriasevia.
Triebelina anterotuberculata[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A bairdiid ostracod, a species of Triebelina.
Triebelina obliquocostata[45]	Sp. nov	Valid	Piovesan, Carbal & Colin in Piovesan et al.	Late Cretaceous (Santonian–Campanian)	Potiguar Basin	Brazil	A bairdiid ostracod, a species of Triebelina.
Triglypta eedemtensis[55]	Sp. nov	Valid	Li in Li et al.	Probably Middle Jurassic	Eedemt Formation	Mongolia	A clam shrimp, a species of Triglypta.
Triglypta yabraiensis[110]	Sp. nov	Valid	Wang			China	A clam shrimp.
Ululapagurus[26]	Gen. et sp. nov	Valid	Fraaije	Late Jurassic (late Kimmeridgian)		Germany	A hermit crab. The type species is Ululapagurus vanbakeli.
Unzhiella navis[111]	Sp. nov	Valid	Voronkova	Late Permian		Russia	A volganellid ostracod, a species of Unzhiella.
Vecticypris samesi[42]	Sp. nov	Valid	Ayress & Whatley	Early Cretaceous		South Atlantic Ocean (North Falkland Basin)	A non-marine ostracod, a species of Vecticypris.
Verruca jagti[67]	Sp. nov	Valid[68]	Gale	Late Cretaceous (Maastrichtian)	Maastricht Formation	Netherlands	A barnacle, a species of Verruca.
Volganella extensis[111]	Sp. nov	Valid	Voronkova	Late Permian		Russia	A volganellid ostracod, a species of Volganella.
Volganella golubevi[111]	Sp. nov	Valid	Voronkova	Late Permian		Russia	A volganellid ostracod, a species of Volganella.
Volganella vjaznikovensis[111]	Sp. nov	Valid	Voronkova	Late Permian		Russia	A volganellid ostracod, a species of Volganella.
Voulteryon[22]	Gen. et sp. nov	Valid	Audo et al.	Middle Jurassic	La Voulte-sur-Rhône Lagerstätte	France	A member of Polychelida. The type species is Voulteryon parvulus.
Wanestheria[38]	Gen. et comb. nov	Valid	Chen & Shen	Late Cretaceous		China	A clam shrimp belonging to the family Sinoestheriidae. Genus includes "Sinoestheria" anhuiensis Chen & Shen (1982), "Sinoestheria" producta Chen & Shen (1982) and "Sinoestheria" semiorbita Chen & Shen (1982).
Youngiverruca[67]	Gen. et sp. nov	Valid[68]	Gale	Late Cretaceous (Maastrichtian)		Germany	A barnacle. The type species is Youngiverruca ruegenensis.
Zovocarcinus[112]	Gen. et sp. nov	Valid	De Angeli & Garassino	Eocene (late Ypresian)		Italy	A member of Panopeidae. The type species is Zovocarcinus muelleri.

Insects

Trilobites

Name	Novelty	Status	Authors	Age	Unit	Location	Notes
Airophrys[113]	Gen. et sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae. The type species is A. balia.
Amphoton zangi[114]	Sp. nov	Valid	Sun, Jago & Bentley	Cambrian (Drumian)	Warburton Basin	Australia	A polymerid trilobite, a species of Amphoton.
Anataphrus kermiti[115]	Sp. nov	Valid	Amati	Late Ordovician	Viola Springs Formation Welling Formation	United States	An isoteline asaphid, a species of Anataphrus.
Anataphrus megalophrys[115]	Sp. nov	Valid	Amati	Late Ordovician	Viola Springs Formation	United States	An isoteline asaphid, a species of Anataphrus.
Asaphellus nazarenensis[116]	Sp. nov	Valid	Tortello & Esteban	Early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Asaphellus.
Asthenopsis conandersoni[117]	Sp. nov	Valid	Bentley & Jago	Cambrian		Australia
Astroproetus franklini[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae.
Bathyurina curtisi[118]	Sp. nov	Valid	Adrain, Karim & Westrop	Ordovician (Floian)	Fillmore Formation	United States	A bathyurid.
Bathyurina hooki[118]	Sp. nov	Valid	Adrain, Karim & Westrop	Ordovician (Floian)	Fillmore Formation	United States	A bathyurid.
Bathyurina morrisi[118]	Sp. nov	Valid	Adrain, Karim & Westrop	Ordovician (Floian)	Fillmore Formation	United States	A bathyurid.
Bathyurina sumneri[118]	Sp. nov	Valid	Adrain, Karim & Westrop	Ordovician (Floian)	Fillmore Formation	United States	A bathyurid.
Bristolia colberti[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Bristolia.
Cambrunicornia saxonica[120]	Sp. nov	Valid	Geyer, Buschmann & Elicki	Middle Cambrian	Tröbitz Formation	Germany	A species of Cambrunicornia.
Caudillaenus[121]	Gen. et sp. nov	Valid	Rábano, Gutiérrez-Marco & García-Bellido	Ordovician (late Darriwilian)	Taddrist Formation	Morocco	An illaenid. The type species is Caudillaenus nicolasi.
Chapmanopyge knudseni[122]	Sp. nov	Valid	McCobb et al.	Early Ordovician		Greenland	A species of Chapmanopyge.
Cheirurus falcatus[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Cheiruridae.
Cybantyx nebulosus[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae.
Cyphaspis heissae[123]	Sp. nov	Valid	Van Viersen & Prescher	Middle Devonian	Ahbach Formation	Germany	An otarionine aulacopleurid, a species of Cyphaspis.
Cyphaspis rommersheimensis[123]	Sp. nov	Valid	Van Viersen & Prescher	Middle Devonian	Freilingen Formation	Germany	An otarionine aulacopleurid, a species of Cyphaspis.
Cyphaspis walteri[123]	Sp. nov	Valid	Van Viersen & Prescher	Middle Devonian	Bou Dib Formation	Morocco	An otarionine aulacopleurid, a species of Cyphaspis.
Daketia[124]	Gen. et sp. nov	Valid	Luo et al.	Ordovician (late Tremadocian-earliest Floian)	Tangchi Formation	China	A member of Asaphida. Genus includes new species D. spinata.
Dalarnepeltis brevifrons[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Tropidocoryphidae.
Diacanthaspis krizi[125]	Sp. nov	Valid	Mergl	Ordovician (late Katian)	Králův Dvůr Formation	Czech Republic	An odontopleurid, a species of Diacanthaspis.
Dicranogmus pearyi[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Lichidae.
Dionide carlottae[126]	Sp. nov	Valid	Corbacho, Morrison & Ait Addi	Ordovician (Katian)	Ktaoua Formation	Morocco	A member of Asaphida belonging to the family Dionididae, a species of Dionide.
Dionidella venusta[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Distyrax bibullatus[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Encrinuridae.
Dorypyge gidgealpaensis[114]	Sp. nov	Valid	Sun, Jago & Bentley	Cambrian (Drumian)	Warburton Basin	Australia	A polymerid trilobite, a species of Dorypyge.
Dulanaspis asper[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Duyunaspis paiwuensis[128]	Sp. nov	Valid	Lei & Peng	Cambrian	Balang Formation	China
Ectenaspis burkhalteri[115]	Sp. nov	Valid	Amati	Late Ordovician	Welling Formation	United States	An isoteline asaphid, a species of Ectenaspis.
Ekwanoscutellum agmen[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae.
Elliptocephala jaredi[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Elliptocephala.
Eoharpes yijianfangensis[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Eokosovopeltis expansus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Erratojincella? decora[129]	Sp. nov	Valid	Yuan & Li	Cambrian	Manto Formation	China	Possibly a species of Erratojincella.
Ganetops[130]	Gen. et comb. nov	Valid	Bignon, Corbacho & López-Soriano	Devonian (late Lochkovian)		Algeria France Germany Spain	A member of Acastidae belonging to the subfamily Asteropyginae. A new genus for "Asteropyge" ebbae R. and E. Richter (1954); genus also contains "Treveropyge" djemelensis Morzadec (1997), "Treveropyge"? cf. ebbae Smeenk (1983), "Paracryphaeus" gerrinensis Timm (1978) and "Acastella" lata Timm (1978).
Geragnostus (Novoagnostus) contractus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Holmiella domackae[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Holmiella.
Holmiella taurus[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Holmiella.
Hungioides yanjinensis[131]	Sp. nov	Valid	Zhou et al.	Ordovician		China	A species of Hungioides.
Hystricurus zanderi[132]	Sp. nov	Valid	Adrain et al.	Ordovician (Tremadocian)		United States
Illaenus cekovioides[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Illaenus tianrongi[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Illaenus usitatus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Iranoleesia (Proasaphiscina?) subconica[129]	Sp. nov	Valid	Yuan & Li	Cambrian	Manto Formation	China	A species of Iranoleesia.
Isotelus bradleyi[115]	Sp. nov	Valid	Amati	Late Ordovician	Viola Springs Formation	United States	An isoteline asaphid, a species of Isotelus.
Isotelus skapaneidos[115]	Sp. nov	Valid	Amati	Late Ordovician	Viola Springs Formation Welling Formation	United States	An isoteline asaphid, a species of Isotelus.
Isotelus violaensis[115]	Sp. nov	Valid	Amati	Late Ordovician	Viola Springs Formation	United States	An isoteline asaphid, a species of Isotelus.
Jungosulcus[113]	Gen. et 3 sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae. The type species is J. emilyae; genus also includes new species J. anaphalantos and J. ventricosus.
Kawina sinica[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China	A species of Kawina[127] or Cydonocephalus.[133]
Kierarges[134]	Gen. et sp. nov	Valid	Corbacho	Ordovician (Floian)	Fezouata Formation Santa Justa Formation	Morocco Portugal	A member of Asaphidae. The type species is Kierarges morrisoni.
Lanceaspis[135]	Gen. et sp. nov	Valid	Corbacho	Devonian (Pragian)	The Mader Platform	Morocco	A member of Acastidae. The type species is Lanceaspis hammondi.
Laneia[113]	Gen. et sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae. The type species is L. enalios.
Ligiscus diana[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae.
Liolalax naresi[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae.
Liostracina tangwangzhaiensis[136]	Sp. nov	Valid	Park et al.	Middle Cambrian		China	A species of Liostracina.
Lisania changi[114]	Sp. nov	Valid	Sun, Jago & Bentley	Cambrian (Drumian)	Warburton Basin	Australia	A polymerid trilobite, a species of Lisania.
Lochmanolenellus pentagonalis[137]	Sp. nov	Valid	Mergl	Cambrian (early Dyeran)		North America	A species of Lochmanolenellus.
Lochmanolenellus subquadratus[137]	Sp. nov	Valid	Mergl	Cambrian (early Dyeran)		North America	A species of Lochmanolenellus.
Lochmanolenellus trapezoidalis[137]	Sp. nov	Valid	Mergl	Cambrian (early Dyeran)		North America	A species of Lochmanolenellus.
Lyralichas qiaoi[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Mesonacis wileyi[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Mesonacis.
Minicryphaeus giganteus[130]	Sp. nov	Valid	Bignon, Corbacho & López-Soriano	Devonian (Pragian)	Ihandar Formation	Morocco	A member of Acastidae belonging to the subfamily Asteropyginae, a species of Minicryphaeus.
Mummaspis delgadoae[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Mummaspis.
Nanillaenus crassus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Nileus amplus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Nileus australis[116]	Sp. nov	Valid	Tortello & Esteban	Early Ordovician (Tremadocian)	Santa Rosita Formation	Argentina	A species of Nileus.
Olenellus baileyi[119]	Sp. nov	Valid	Gapp & Lieberman	Early Cambrian	Sekwi Formation	Canada	A member of Olenelloidea, a species of Olenellus.
Olenoides sagittatus[138]	Sp. nov	Valid	Geyer in Geyer et al.	Cambrian (Amgan)		Kyrgyzstan	A dorypygid corynexochidan, a species of Olenoides.
Opoa limatula[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Scutelluidae.
Owensus[113]	Gen. et sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae. The type species is O. arktoperates.
Parakoldinioidia akerfeldti[139]	Sp. nov	Valid	Westrop & Adrain	Cambrian	Signal Mountain Formation	United States	A member of Missisquoiidae.
Parakoldinioidia akessoni[139]	Sp. nov	Valid	Westrop & Adrain	Cambrian	Signal Mountain Formation	United States	A member of Missisquoiidae.
Parakoldinioidia lindgreni[139]	Sp. nov	Valid	Westrop & Adrain	Cambrian	Signal Mountain Formation Wilberns Formation	United States	A member of Missisquoiidae.
Parakoldinioidia lopezi[139]	Sp. nov	Valid	Westrop & Adrain	Cambrian	Signal Mountain Formation	United States	A member of Missisquoiidae.
Parakoldinioidia mendezi[139]	Sp. nov	Valid	Westrop & Adrain	Cambrian	Signal Mountain Formation	United States	A member of Missisquoiidae.
Paraperaspis[127]	Gen. et 2 sp. et comb. nov	Valid	Zhou, Yin & Zhou	Ordovician		China	A nileid. Genus includes new species P. angustus and P. dawangouensis, as well as "Nileus" convergens Lu (1975).
Paraplethopeltis helli[132]	Sp. nov	Valid	Adrain et al.	Ordovician (Tremadocian)		United States
Parillaenus exiguus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Parillaenus vetus[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Perryus mikulici[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Encrinuridae.
Profallotaspis tyusserica[140]	Sp. nov	Valid	Bushuev & Goryaeva in Bushuev, Goryaeva & Pereladov	Cambrian	Tyusser Formation	Russia	A member of Fallotaspididae.
Proetus confluens[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae.
Protolenus (Hupeolenus) bergstroemi[120]	Sp. nov	Valid	Geyer, Buschmann & Elicki	Middle Cambrian	Tröbitz Formation	Germany	A species of Protolenus.
Pseudinouyia shiliuyuanensis[129]	Sp. nov	Valid	Yuan & Li	Cambrian	Manto Formation	China	A species of Pseudinouyia.
Pseudoclelandia weymouthae[132]	Sp. nov	Valid	Adrain et al.	Ordovician (Tremadocian)		United States
Pseudohystricurus wigglesorum[141]	Sp. nov	Valid	Adrain, Karim & Westrop	Ordovician	Garden City Formation	United States	A dimeropygid trilobite.
Ptychobaba[142]	Gen. et comb. nov	Valid	Pratt & Bordonaro	Cambrian	La Laja Formation Rome Formation	Argentina United States	A new genus for "Ptychoparella" buttsi Resser (1938).
Punka adamsi[122]	Sp. nov	Valid	McCobb et al.	Early Ordovician		Greenland	A species of Punka.
Radiurus pauli[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Cheiruridae.
Raragnostus shergoldi[114]	Sp. nov	Valid	Sun, Jago & Bentley	Cambrian (Drumian)	Warburton Basin	Australia	An agnostoid trilobite, a species of Raragnostus.
Rossaspis leboni[132]	Sp. nov	Valid	Adrain et al.	Ordovician (Tremadocian)		United States
Sagavia laxa[127]	Sp. nov	Valid	Zhou, Yin & Zhou	Ordovician		China
Sestrostega cylindrica[129]	Sp. nov	Valid	Yuan & Li	Cambrian	Manto Formation	China	A species of Sestrostega.
Shanxiella (Shanxiella) latilimbata[129]	Sp. nov	Valid	Yuan & Li	Cambrian	Manto Formation	China	A species of Shanxiella.
Solenoparia gatehousei[114]	Sp. nov	Valid	Sun, Jago & Bentley	Cambrian (Drumian)	Warburton Basin	Australia	A polymerid trilobite, a species of Solenoparia.
Stegnopsis erythragora[115]	Sp. nov	Valid	Amati	Late Ordovician	Viola Springs Formation Welling Formation	United States	An isoteline asaphid, a species of Stegnopsis.
Stegnopsis wellingensis[115]	Sp. nov	Valid	Amati	Late Ordovician	Welling Formation	United States	An isoteline asaphid, a species of Stegnopsis.
Stenopareia persica[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Illaenidae.
Struveaspis bignoni[143]	Sp. nov	Valid	Corbacho	Devonian (Eifelian)		Morocco	A member of Phacopidae, a species of Struveaspis.
Tairongia[127]	Gen. et sp. et comb. nov	Valid	Zhou, Yin & Zhou	Ordovician		China	A lichid. Genus includes new species T. bachuensis, as well as "Lichas" browni Sun (1931).
Trinodus jii[127]	Nom. nov		Zhou, Yin & Zhou	Ordovician		China	A replacement name for Trinodus latilimbatus Ji (1986).
Winiskia eruga[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae.
Winiskia leptomedia[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae.
Winiskia stickta[113]	Sp. nov	Valid	Hughes & Thomas	Silurian		Greenland	A member of the family Proetidae.

Other arthropods

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Anbarrhacus[144]	Gen. et sp. nov	Valid	Riquelme & Hernández in Riquelme et al.	Early or middle Miocene		Mexico	A platyrhacid polydesmidan millipede. The type species is Anbarrhacus adamantis.	Anbarrhacus adamantis
Annge[145]	Gen. et sp. nov	Valid	Smith et al.	Cambrian (Templetonian)	Giles Creek Dolostone	Australia	A member of Bradoriida. Genus includes new species A. iperte.
Cambrolongispina[146]	Gen. et 2 sp. nov	Valid	Zhang, Dong & Xiao	Cambrian (Drumian)		China	A bivalved arthropod of uncertain phylogenetic placement. Genus contains two species: Cambrolongispina reticulata and Cambrolongispina glabra.
Erjiecaris[147]	Gen. et sp. nov	Valid	Fu, Zhang & Budd	Early Cambrian	Chengjiang Lagerstätte	China	An early bivalved arthropod of uncertain phylogenetic placement. The type species is Erjiecaris minusculo.
Falcatamacaris[148]	Gen. et sp. nov	Valid[149]	Ortega-Hernández et al.	Cambrian (Guzhangian)	Weeks Formation	United States	An artiopodan arthropod. The type species is Falcatamacaris bellua.
Flemingopsis anteriospinata[145]	Sp. nov	Valid	Smith et al.	Cambrian (Templetonian)	Giles Creek Dolostone	Australia	A member of Bradoriida.
Haifengella[150]	Gen. et sp. nov	Valid	Zhao et al.	Early Cambrian	Chengjiang Lagerstätte	China	An arthropod related to Helmetia. The type species is Haifengella corona.
Jiucunella phaseloa[151]	Sp. nov	Valid	Betts et al.	Early Cambrian		Australia	A member of Bradoriida, a species of Jiucunella.
Jixinlingella daimonikoa[151]	Sp. nov	Valid	Betts et al.	Early Cambrian		Australia	A member of Bradoriida, a species of Jixinlingella.
Limulus darwini[152]	Sp. nov	Valid	Kin & Błażejowski	Late Jurassic (late Tithonian)	Kcynia Formation	Poland	A horseshoe crab. Originally described as a species of Limulus; Bicknell et al. (2021) transferred this species to the genus Crenatolimulus.[153]
Maatidesmus[144]	Gen. et sp. nov	Valid	Riquelme & Hernández in Riquelme et al.	Early or middle Miocene (c. 23–15 Mya)	Mexican amber	Mexico	A chelodesmid polydesmidan millipede. The type species is M. paachtun.	Maatidesmus paachtun
Monasterium ferox[145]	Sp. nov	Valid	Smith et al.	Cambrian (Templetonian)	Giles Creek Dolostone	Australia	A member of Bradoriida.
Mongolitubulus anthelios[151]	Sp. nov	Valid	Betts et al.	Early Cambrian		Australia	A member of Bradoriida, a species of Mongolitubulus.
Mongolitubulus tunpere[145]	Sp. nov	Valid	Smith et al.	Cambrian (Templetonian)	Giles Creek Dolostone	Australia	A member of Bradoriida.
Neokunmingella moroensis[151]	Sp. nov	Valid	Betts et al.	Early Cambrian		Australia	A member of Bradoriida, a species of Neokunmingella.
Polzia[154]	Gen. et sp. nov	Valid	Hegna, Vega & González-Rodríguez	Cretaceous (Albian to Cenomanian)	El Doctor Formation	Mexico	A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans). The type species is Polzia eldoctorensis.
Sinskolutella cuspidata[151]	Sp. nov	Valid	Betts et al.	Early Cambrian		Australia	A member of Bradoriida, a species of Sinskolutella.
Thylacares[155]	Gen. et sp. nov	Valid	Haug et al.	Silurian (late Telychian)	Brandon Bridge Formation	United States	A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans). The type species is Thylacares brandonensis.
Victoriacaris[154]	Gen. et sp. nov	Valid	Hegna, Vega & González-Rodríguez	Cretaceous (Albian to Cenomanian)	El Doctor Formation	Mexico	A member of Thylacocephala (a group of arthropods of uncertain phylogenetic placement, possibly crustaceans. The type species is Victoriacaris muhiensis.

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[47] Matúš Hýžný; Martina Kočová Veselská; Pavel Dvořák (2014). "On the occurrence of Ctenocheles (Decapoda, Axiidea, Ctenochelidae) in the Bohemian Cretaceous Basin". Bulletin of Geosciences. 89 (2): 245–256. doi:10.3140/bull.geosci.1421. PMC 4430802. PMID 25983568.

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[125] Michal Mergl (2014). "Odontopleurid trilobites of the Katian/Hirnantian boundary interval in the Prague Basin (Bohemia)". Bulletin of Geosciences. 89 (2): 401–412. doi:10.3140/bull.geosci.1452.

[126] Joan Corbacho; Scott Morrison; Abdellah Ait Addi (2014). "Dionide carlottae: Una nueva especie de Dionididae (Trilobita) del Ordovícico superior de Marruecos". Batalleria. 21: 13–21.

[Paraperaspis-127] Zhiyi Zhou; Gongzheng Yin; Zhiqiang Zhou (2014). "Ordovician (Darriwilian-early Katian) trilobite faunas of northwestern Tarim, Xinjiang, China". Memoirs of the Association of Australasian Palaeontologists. 46: 1–142.

[128] Qian-ping Lei; Shan-chi Peng (2014). "Duyunaspis Zhang and Qian in Zhou et al., 1977 (Trilobita) from the Balang Formation (Cambrian, Qiandongian) in northwestern Hunan Province and its intraspecific variations". Acta Palaeontologica Sinica. 53 (3): 352–362. doi:10.19800/j.cnki.aps.2014.03.010.

[APJ534497-129] Jin-liang Yuan; Yue Li (2014). "Trilobites of the uppermost part of the Manto Formation (Hsuchuangian) at the Shiliuyuan, Yicheng District, Zaozhuang City, Shandong". Acta Palaeontologica Sinica. 53 (4): 497–526. doi:10.19800/j.cnki.aps.2014.04.012.

[Ganetops-130] Arnaud Bignon; Joan Corbacho; Francisco J. López-Soriano (2014). "A revision of the first Asteropyginae (Trilobita; Devonian)". Geobios. 47 (5): 281–289. doi:10.1016/j.geobios.2014.09.002.

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[IbexianSeries-132] Jonathan M. Adrain; Stephen R. Westrop; Talia S. Karim; Ed Landing (2014). "Trilobite biostratigraphy of the Stairsian Stage (upper Tremadocian) of the Ibexian Series, Lower Ordovician, western United States". Memoirs of the Association of Australasian Palaeontologists. 45: 167–214.

[133] Jonathan M. Adrain; Francesc Pérez-Peris (2021). "Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada". Zootaxa. 5041 (1): 1–73. doi:10.11646/zootaxa.5041.1.1. PMID 34811019. S2CID 239237412.

[134] Joan Corbacho (2014). "Kierarges morrisoni: Un nuevo género de Isotelinae (Trilobita) del Floiense de Portugal y Marruecos". Batalleria. 20: 21–26.

[135] Joan Corbacho (2014). "Lanceaspis hammondi n. gen., n. sp. – a questionable early Asteropyginae (Trilobita) from the Pragian of the Morocco". Batalleria. 20: 3–10.

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[ZTLochmanolenellus-137] Mark Webster; Lisa L. Bohach (2014). "Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia". Zootaxa. 3824 (1): 1–66. doi:10.11646/zootaxa.3824.1.1. PMID 24989880.

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[Memoirs45Parakoldinioidia-139] Stephen R. Westrop; Jonathan M. Adrain (2014). "The missisquoiid trilobite Parakoldinioidia Endo 1937 in the uppermost Cambrian of Oklahoma and Texas, and its biostratigraphic significance". Memoirs of the Association of Australasian Palaeontologists. 45: 117–152.

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[142] Brian R. Pratt; Osvaldo L. Bordonaro (2014). "Early middle Cambrian trilobites from La Laja Formation, Cerro El Molle, Precordillera of western Argentina". Journal of Paleontology. 88 (5): 906–924. doi:10.1666/13-083. S2CID 131408281.

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[Anbarrhacus-144] Francisco Riquelme; Miguel Hernández-Patricio; Arnulfo Martínez-Dávalos; Mercedes Rodríguez-Villafuerte; Maira Montejo-Cruz; Jesús Alvarado-Ortega; José L. Ruvalcaba-Sil; Luis Zúñiga-Mijangos (2014). "Two Flat-Backed Polydesmidan Millipedes from the Miocene Chiapas-Amber Lagerstätte, Mexico". PLOS ONE. 9 (8): e105877. Bibcode:2014PLoSO...9j5877R. doi:10.1371/journal.pone.0105877. PMC 4146559. PMID 25162220.

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[146] Huaqiao Zhang; Xi-ping Dong; Shuhai Xiao (2014). "New Bivalved Arthropods from the Cambrian (Series 3, Drumian Stage) of Western Hunan, South China". Acta Geologica Sinica (English Edition). 88 (5): 1388–1396. doi:10.1111/1755-6724.12306. S2CID 129703700.

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