2016 in paleoichthyology

This list of fossil fishes described in 2016 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes and other fishes of every kind that have been described during the year 2016, as well as other significant discoveries and events related to paleontology of fishes that occurred in the year 2016. The list only includes taxa at the level of genus or species.

List of years in paleoichthyology
In paleontology
2013
2014
2015
2016
2017
2018
2019
In science
2013
2014
2015
2016
2017
2018
2019
In paleobotany
2013
2014
2015
2016
2017
2018
2019
In arthropod paleontology
2013
2014
2015
2016
2017
2018
2019
In paleoentomology
2013
2014
2015
2016
2017
2018
2019
In paleomalacology
2013
2014
2015
2016
2017
2018
2019
In reptile paleontology
2013
2014
2015
2016
2017
2018
2019
In archosaur paleontology
2013
2014
2015
2016
2017
2018
2019
In mammal paleontology
2013
2014
2015
2016
2017
2018
2019

Research

  • Tullimonstrum gregarium is argued to be a stem-lamprey by McCoy et al. (2016).[1]
  • A study of the eye anatomy of Tullimonstrum gregarium is published by Clements et al. (2016).[2]
  • New information on the anatomy of the Carboniferous taxa Mayomyzon pieckoensis and Myxinikela siroka is published by Gabbott et al. (2016), who report the presence of melanosomes in their eyes; the authors also conduct a study on the phylogenetic relationships of these species, finding Mayomyzon to be a relative of lampreys and finding Myxinikela to be a relative of hagfish.[3]
  • A study on the phylogenetic relationships of Palaeospondylus gunni is published by Hirasawa, Oisi & Kuratani (2016), who interpret it as a probable stem-hagfish.[4]
  • New information on the dermal bone histology and phylogenetic relationships of the jawless anaspids, published by Keating & Donoghue (2016), reveal that they are a monophyletic group nested within skeletonizing vertebrates, rather than early relatives of lampreys and hagfish.[5]
  • A study on the phylogenetic relationships of early jawed vertebrates is published by Qiao et al. (2016).[6]
  • A study on the ontogenetic composition of the Devonian placoderm material known from the Strud quarry (Namur Province, Belgium) is published by Olive et al. (2016), who interpret the Strud locality as representing a placoderm nursery.[7]
  • A description of the general anatomy, morphology and histology of dermal elements and the endoskeletal shoulder girdle of three Devonian (Eifelian-Givetian) species of acanthodians known from Scotland (Diplacanthus crassisimus, Diplacanthus tenuistriatus and Rhadinacanthus longispinus) is published by Burrow et al. (2016).[8]
  • Fossil tooth of a member of the genus Lagarodus (a cartilaginous fish belonging to the group Euchondrocephali, of uncertain phylogenetic placement within the latter group) is reported for the first time from the Carboniferous of Svalbard by Cuny, Kristensen & Stemmerik (2016).[9]
  • A study on the enameloid ultrastructure of the teeth of members of the genus Ptychodus recovered from the Lincoln Limestone of the Greenhorn Formation (Barton County, Kansas, USA) and its implications for the phylogenetic placement of the genus is published by Hoffman, Hageman & Claycomb (2016).[10]
  • A specimen of Galeorhinus cuvieri (fossil relative of the school shark) with preserved soft tissues and stomach contents (indicating that it preyed on Eocene relatives of the living barracudas) is described from the Eocene (Ypresian) Monte Bolca site in Italy by Fanti et al. (2016).[11]
  • A redescription of the Miocene shark Glyphis pagoda is published by Shimada et al. (2016).[12]
  • A study of geographical distribution patterns and global abundance of Carcharocles megalodon from the Miocene to the Pliocene, and a discussion of the possible causes of its extinction, is published by Pimiento et al. (2016).[13]
  • A study on the growth history of the teeth of Andreolepis hedei is published by Chen et al. (2016).[14]
  • A study on the anatomical diversification of teleosts and holosteans during 160 million years of their evolution (PermianEarly Cretaceous) is published by Clarke, Lloyd & Friedman (2016).[15]
  • Specimens of Saurichthys costasquamosus, Saurichthys macrocephalus and Saurichthys paucitrichus with preserved casts of gastrointestinal tract are described by Argyriou et al. (2016).[16]
  • A redescription of Ionoscopus petrarojae and a study on the phylogenetic relationships of the species is published by Taverne & Capasso (2016).[17]
  • A redescription and a study on the phylogenetic relationships of Dapedium pholidotum, based on new fossil material from the Jurassic (Toarcian) Posidonia Shale (Germany) is published by Thies & Waschkewitz (2016), who name a new order Dapediiformes including the family Dapediidae.[18]
  • A redescription and a study on the phylogenetic relationships of the dapediiform species Hemicalypterus weiri is published by Gibson (2016).[19]
  • A study on the anatomy of the Carboniferous teleost relative Aetheretmon and its implications for the evolution of tail and caudal fin of vertebrates in published by Sallan (2016).[20]
  • A redescription of the anatomy of the Early Cretaceous osteoglossiform Chanopsis lombardi known from the Democratic Republic of the Congo is published by Taverne (2016).[21]
  • A specimen of an osteoglossid fish, putatively referred to the species Ridewoodichthys caheni, is described from the Paleocene (Danian) of Angola by Taverne (2016).[22]
  • A study on the anatomy and phylogenetic relationships of the ellimmichthyiform Codoichthys carnavalii is published by de Figueiredo & Ribeiro (2016).[23]
  • Fossilized hearts are reported in two specimens of Rhacolepis buccalis by Maldanis et al. (2016).[24]
  • A phylogenetic study of spiny-rayed teleosts, including Late Cretaceous fossil taxa and that recovers the molecular-based interrelationships with morphological information for the first time, is published by Davesne et al. (2016).[25]
  • Meemannia eos, initially classified as a lobe-finned fish, is reinterpreted as an early-diverging ray-finned fish by Lu et al. (2016).[26]
  • Description of new skull material of Qingmenodus yui from Pragian of China and a study of phylogenetic relationships of onychodonts is published by Lu et al. (2016).[27]
  • Clement et al. (2016) reconstruct the shape of the brain of the Devonian lungfish Rhinodipterus kimberleyensis on the basis of a CT scan of its endocast.[28]
  • Virtual cranial endocast of Dipnorhynchus sussmilchi is reconstructed by Clement et al. (2016).[29]

New taxa

Jawless vertebrates

Name Novelty Status Authors Age Unit Location Notes Images

Arianalepis[30]

Gen. et sp. nov

Valid

Hairapetian et al.

Devonian (Famennian)

 Australia
 Iran

A turiniid thelodont. The type species is Arianalepis megacostata; genus also contains a second, indeterminate species.

Kodinskaspis[31]

Gen. et sp. nov

Valid

Dzik & Moskalenko

Ordovician (460 Mya)

Mamyry Formation

 Russia

A possible relative of Tesakoviaspis concentrica. The type species is Kodinskaspis angarensis.

Neoturinia[32]

Gen. et comb. nov

Valid

Hairapetian, Blom & Turner

Devonian (Frasnian)

 Iran

A turiniid thelodont. The type species is "Turinia" hutkensis Blieck & Goujet (1978).

Phyllonaspis[33]

Gen. et 3 sp. nov

Valid

Elliott

Early Devonian (late Emsian)

Lost Burro Formation
Sevy Dolomite

 United States
( California
 Nevada
 Utah)

A member of Cyathaspididae. The type species is P. laevis; genus also includes P. serratus and P. taphensis.

Placoderms

Name Novelty Status Authors Age Unit Location Notes Images

Bothriolepis rex[34]

Sp. nov

Valid

Downs et al.

Devonian (Frasnian)

Nordstrand Point Formation

 Canada
( Nunavut)

A member of Antiarchi.

Qilinyu[35]

Gen. et sp. nov

Valid

Zhu et al.

Silurian (late Ludlow)

Kuanti Formation

 China

The type species is Q. rostrata.

Acanthodians

Name Novelty Status Authors Age Unit Location Notes Images

Atopacanthus juvai[36]

Sp. nov

Valid

Hairapetian & Burrow

Late Devonian

Bahram Formation

 Iran

A member of Ischnacanthiformes; a species of Atopacanthus.

Cartilaginous fishes

Name Novelty Status Authors Age Unit Location Notes Images

Akaimia myriacuspis[37]

Sp. nov

Valid

Srdic, Duffin & Martill

Middle Jurassic (Callovian)

Peterborough Member of the Oxford Clay Formation

 United Kingdom

A carpet shark.

Ceolometlaouia[38]

Gen. et sp. nov

Valid

Engelbrecht et al.

Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A carpet shark. Genus includes new species C. pannucae.

Crassodus[39]

Gen. et sp. nov

Valid

Maisch & Matzke

Early Jurassic (early Toarcian)

Posidonienschiefer Formation

 Germany

A member of Hybodontidae. The type species is C. reifi.

Deltalepis[40]

Gen. et 2 sp. nov

Valid

Andreev et al.

Silurian

Chargat Formation

 Mongolia

A relative of Elegestolepis and Ellesmereia. Genus includes new species D. magna and D. parva.

Haimirichia[41]

Gen. et comb. nov

Valid[42]

Vullo, Guinot & Barbe

Cretaceous

 Angola
 Egypt
 France
 Japan
 Morocco
 Nigeria
 Spain
 United States

A member of Lamniformes of uncertain phylogenetic placement; a new genus for "Odontaspis" amonensis Cappetta & Case (1975).

Isanodus nongbualamphuensis[43]

Sp. nov

Valid

Khamha, Cuny & Lauprasert

Early Cretaceous

 Thailand

A member of Hybodontiformes, a species of Isanodus.

Lonchidion derenzii[44]

Sp. nov

Valid

Manzanares et al.

Late Triassic

 Spain

A member of Hybodontiformes.

Megachasma alisonae[45]

Sp. nov

Valid

Shimada & Ward

Late Eocene

Søvind Marl Formation

 Denmark

A relative of the megamouth shark.

Megalolamna[46]

Gen. et sp. nov

Valid

Shimada et al.

Miocene (AquitanianBurdigalian)

Chilcatay Formation
Jewett Sand Formation
Oi Formation
O'oshimojo Formation
Pungo River Formation
Uitpa Formation

 Colombia
 Japan
 Peru
 United States
( California,
 North Carolina)

A member of Otodontidae. The type species is M. paradoxodon.

Notoramphoscyllium[38]

Gen. et sp. nov

Valid

Engelbrecht et al.

Eocene

La Meseta Formation

Antarctica
(Seymour Island)

A carpet shark. Genus includes new species N. woodwardi.

Oligodalatias[47]

Gen. et sp. nov

Valid

Welton

Early Eocene-Miocene

Keasey Formation
Kirker Sandstone
Lillebælt Clay
Pittsburg Bluff Formation

 Denmark
 France
 United States
( California
 Oregon)

A member of Dalatiidae. The type species is O. jordani.

Orectolobus ziegenhinei[48]

Sp. nov

Valid

Cappetta & Case

Eocene (Lutetian)

Lisbon Formation

 United States
( Alabama)

A species of Orectolobus.

Orthechinorhinus davidae[49]

Sp. nov

Valid

Welton

Early Oligocene

Alsea Formation

 United States
( Oregon)

Probably a member of Etmopteridae.

Ossianodus[50]

Gen. et sp. nov

Valid

Ginter

Carboniferous (Gzhelian)

Indian Cave Sandstone

 United States
( Nebraska)

A member of Hybodontidae. The type species is O. nebraskensis.

Pristiophorus laevis[51]

Sp. nov

Valid

Engelbrecht et al.

Eocene

Antarctica

A species of Pristiophorus.

Protosqualus argentinensis[52]

Sp. nov

Valid

Bogan, Agnolin & Novas

Late Cretaceous (Maastrichtian)

Calafate Formation

 Argentina

A member of Squaliformes, a species of Protosqualus.

Rhinoscymnus viridiadamas[53]

Sp. nov

Valid

Welton & Goedert

Oligocene

Lincoln Creek Formation

 United States
( Washington)

A member of Somniosidae, a species of Rhinoscymnus.

Scoliodon conecuhensis[48]

Sp. nov

Valid

Cappetta & Case

Eocene (Lutetian)

Lisbon Formation

 United States
( Alabama)

A species of Scoliodon.

Solinalepis[54]

Gen. et sp. nov

Valid

Andreev et al.

Ordovician (Sandbian)

Harding Sandstone

 United States
( Colorado)

A member of Mongolepidida of uncertain phylogenetic placement. The type species is S. levis.

Somniosus gonzalezi[53]

Sp. nov

Valid

Welton & Goedert

Oligocene

Pysht Formation

 United States
( Washington)

A member of Somniosidae, a species of Somniosus.

Sphenacanthus tenuis[50]

Sp. nov

Valid

Ginter

Carboniferous (Gzhelian)

Indian Cave Sandstone

 United States
( Nebraska)

A member of Sphenacanthidae.

Squalicorax deckeri[55]

Sp. nov

Valid

Bice & Shimada

Late Cretaceous (Turonian)

Codell Sandstone Member of the Carlile Shale

 United States
( Kansas)

Squatina (Squatina) fortemordeo[56]

Sp. nov

Valid

Siversson et al.

Late Cretaceous (early Campanian)

Kristianstad Basin

 Sweden

An angel shark.

Squatina (Squatina) lundegreni[56]

Sp. nov

Valid

Siversson et al.

Late Cretaceous (early Campanian)

Kristianstad Basin

 Sweden

An angel shark.

Synechodus filipi[56]

Sp. nov

Valid

Siversson et al.

Late Cretaceous (early Campanian)

Kristianstad Basin

 Sweden

A member of Synechodontiformes, a species of Synechodus.

Tethylamna[48]

Gen. et sp. et comb. nov

Valid

Cappetta & Case

Eocene (Lutetian-Priabonian)

Lisbon Formation

 Egypt
 Morocco
 Pakistan
 United States
( Alabama
 Georgia (U.S. state))

Genus includes new species T. dunni, as well as "Lamna" twiggsensis Case (1981).

Bony fishes

Name Novelty Status Authors Age Unit Location Notes Images

Adrianaichthys[57]

Gen. et comb. nov

Valid

Meunier et al.

Late Cretaceous (Cenomanian)

Kem Kem Group

 Morocco

A member of Lepisosteiformes; a new genus for "Lepidotes" pankowskii Forey, López-Arbarello & MacLeod (2011).

Altamuraichthys[58]

Gen. et sp. nov

Valid

Taverne

Late Cretaceous (Campanian-Maastrichtian)

 Italy

A member of Ichthyodectidae. The type species is A. meleleoi.

Amazonasciaena[59]

Gen. et comb. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Aquitanian to early Burdigalian)

Pirabas Formation

 Brazil

A member of Sciaenidae; a new genus for "Sciaenops" rossettiae Aguilera & Schwarzhans (2014).

Amyzon kishenehnicum[60]

Sp. nov

Valid

Liu, Wilson & Murray

Eocene

Kishenehn Formation

 United States
( Montana)

A member of Catostomidae.

Argentina antarctica[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A herring smelt, a species of Argentina.

Ariomma? astridae[62]

Sp. nov

Valid

Hoedemakers & Schneider

Oligocene (Rupelian)

 Germany

Possibly a species of Ariomma.

Armigatus dalmaticus[63]

Sp. nov

Valid

Murray et al.

Late Cretaceous (Campanian)

 Croatia

A clupeomorph belonging to the group Ellimmichthyiformes.

Askerichthys[64]

Gen. et sp. nov

Valid

Borgen & Nakrem

Late Carboniferous

Tanum Formation

 Norway

A member of the family Megalichthyidae. The type species is A. heintzi.

Atractoscion odeai[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Burdigalian to early Langhian)

Cantaure Formation
Castilletes Formation

 Colombia
 Venezuela

A member of Sciaenidae, a species of Atractoscion

Axelrodichthys megadromos[65]

Sp. nov

Valid

Cavin, Valentin & Garcia

Late Cretaceous (early Campanian)

 France

A mawsoniid coelacanth, a species of Axelrodichthys.

Balkaria[66]

Gen. et sp. nov

Valid[67]

Bannikov et al.

Eocene

 Russia
( Kabardino-Balkaria)

A member of Tetraodontiformes belonging to the suborder Tetraodontoidei. The type species is Balkaria histiopterygia.

‘Branchiostegus’ brevis[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Branchiostegus

Callaus cubaguanus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Miocene to late Pliocene

Cubagua Formation

 Venezuela

A member of Sciaenidae, a species of Callaus

Carlomonnius[70]

Gen. et sp. nov

Valid

Bannikov & Carnevale

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of Gobioidei of uncertain phylogenetic placement. The type species is Carlomonnius quasigobius.

Cataetyx stringeri[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Late Pliocene to early Pleistocene

Bowden Formation
Moin Formation

 Costa Rica
 Jamaica

A viviparous brotula; a species of Cataetyx.

Catutoichthys[72]

Gen. et sp. nov

Valid

Gouiric-Cavalli

Late Jurassic (Tithonian)

Vaca Muerta Formation

 Argentina

A member of Caturidae. The type species is C. olsacheri.

Cepola anderssoni[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A bandfish, a species of Cepola.

Cepola rostislavi[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Cepola

Ceratodus kranzi[73]

Sp. nov

Valid

Frederickson, Lipka & Cifelli

Early Cretaceous

Potomac Formation

 United States
( Maryland)

A lungfish, a species of Ceratodus.

Coelorinchus balushkini[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A grenadier, a species of Coelorinchus.

Coelorinchus nordenskjoeldi[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A grenadier, a species of Coelorinchus.

Ctenoplectus[74]

Gen. et sp. nov

Valid

Close et al.

Eocene (Ypresian)

London Clay Formation

 United Kingdom

A member of Tetraodontiformes related to the threetooth puffer. The type species is C. williamsi.

Cumbaaichthys[75]

Gen. et sp. nov

Valid

Murray

Late Cretaceous (Turonian)

 Canada

A member of Polymixiiformes of uncertain phylogenetic placement. The type species is Cumbaaichthys oxyrhynchus.

Cynoscion latiostialis[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian)

Gatun Formation
Urumaco Formation

 Panama
 Venezuela

A member of Sciaenidae, a species of Cynoscion

Cynoscion prolixus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian) to late Pliocene

Chagres Formation
Cubagua Formation
Urumaco Formation

 Panama
 Venezuela

A member of Sciaenidae, a species of Cynoscion

Cynoscion scitulus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian)

Gatun Formation

 Panama

A member of Sciaenidae, a species of Cynoscion

Diaphus? marambionis[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A lanternfish, possibly a species of Diaphus.

Ebertichthys[76]

Gen. et sp. nov

Valid

Arratia

Late Jurassic

 Germany

A teleost related to Ascalabos voithii. The type species is Ebertichthys ettlingensis.

Eekaulostomus[77]

Gen. et sp. nov

Valid

Cantalice & Alvarado-Ortega

Paleocene (Danian)

 Mexico

A member of the superfamily Aulostomoidea. The type species is E. cuevasae.

‘Elops’ miiformis[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Elops

Eoengraulis[78]

Gen. et sp. nov

Valid

Marramà & Carnevale

Eocene (late Ypresian)

Monte Bolca locality

 Italy

An anchovy. The type species is Eoengraulis fasoloi.

‘Evermannella’ razza[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Evermannella

Foreyclupea[79]

Gen. et sp. nov

Valid

Vernygora, Murray & Wilson

Early Cretaceous (Albian)

Loon River Formation

 Canada
( Northwest Territories)

A member of Clupeomorpha, possibly related to Ranulfoichthys dorsonudum and Scutatuspinosus itapagipensis. The type species is Foreyclupea loonensis.

Francolebias arvernensis[80]

Sp. nov

Valid

Gaudant

Oligocene

 France

A relative of Valencia.

Frodoichthys[81]

Gen. et sp. nov

Valid

Sun et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A member of Neopterygii of uncertain phylogenetic placement, showing similarities to Prosantichthys and thus to the halecomorph order Panxianichthyformes. The type species is F. luopingensis.

Gimlichthys[81]

Gen. et sp. nov

Valid

Sun et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A member of Neopterygii of uncertain phylogenetic placement, more similar to ginglymodians than to halecomorphs. The type species is G. dawaziensis.

Gladiopycnodus byrnei[82]

Sp. nov

Valid

Marramà et al.

Late Cretaceous (Cenomanian)

 Lebanon

A member of Pycnodontiformes belonging to the superfamily Coccodontoidea, a species of Gladiopycnodus.

Habroichthys dolomiticus[83]

Sp. nov

Valid

Tintori, Lombardo & Kustatscher

Middle Triassic (Anisian)

 Italy

‘Haemulon’ strascinate[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Haemulon

Herreraichthys[84]

Gen. et sp. nov

Valid

Alvarado-Ortega et al.

Late Cretaceous (Santonian)

 Mexico

A gar. The type species is Herreraichthys coahuilaensis.

Hoplobrotula? antipoda[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A cusk-eel, possibly a species of Hoplobrotula.

Hoplopteryx dakotaensis[85]

Sp. nov

Valid

Grandstaff & Parris

Late Cretaceous (Turonian)

Carlile Formation

 United States
( South Dakota)

Hyporhamphus tatjanchenkoi[86]

Sp. nov

Valid

Bannikov, Carnevale & Kotlyar

Miocene

 Russia
( Krasnodar Krai)

A halfbeak, a species of Hyporhamphus.

Igornichthys bohemicus[87]

Sp. nov

Valid

Štamberg

Permian (Asselian)

Vrchlabí Formation

 Czech Republic

A ray-finned fish belonging to the group Brookvaliiformes.

Isopisthus acer[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Pliocene

Río Banano Formation

 Costa Rica

A member of Sciaenidae, a species of Isopisthus

Khoratichthys[88]

Gen. et sp. nov

Valid

Deesri, Jintasakul & Cavin

Late Jurassic–Early Cretaceous

Phu Kradung Formation

 Thailand

A basal member of Lepisosteiformes. The type species is K. gibbus.

Kooiichthys[89]

Gen. et sp. nov

Valid

Azpelicueta et al.

Miocene

Puerto Madryn Formation

 Argentina

A catfish. The type species is Kooiichthys jono.

Lamprogrammus manzanilla[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (late Tortonian)

Manzanilla Formation

 Trinidad and Tobago
(Trinidad)

A cusk-eel; a species of Lamprogrammus.

Larimus angosturae[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian) to Pliocene (Zanclean)

Angostura Formation
Río Banano Formation

 Costa Rica
 Ecuador

A member of Sciaenidae, a species of Larimus

Larimus humboldti[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Burdigalian to early Langhian)

Cantaure Formation
Jimol Formation

 Colombia
 Venezuela

A member of Sciaenidae, a species of Larimus

Larimus pandus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian)

Angostura Formation
Gatun Formation

 Ecuador
 Panama

A member of Sciaenidae, a species of Larimus

Lepophidium borbonensis[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (Tortonian and Messinian)

Angostura Formation
Onzole Formation

 Ecuador

A cusk-eel; a species of Lepophidium.

Lepophidium crebrum[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (Serravallian and Tortonian)

Angostura Formation
Manzanilla Formation
Tamana Formation

 Ecuador
 Trinidad and Tobago
(Trinidad)

A cusk-eel; a species of Lepophidium.

Lepophidium gentilis[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (Tortonian)

Gatun Formation

 Panama

A cusk-eel; a species of Lepophidium.

Lepophidium leai[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Late Pliocene

Cubagua Formation

 Venezuela

A cusk-eel; a species of Lepophidium.

Lepophidium limulum[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (Messinian)

Onzole Formation

 Ecuador

A cusk-eel; a species of Lepophidium.

Lepophidium refugum[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Early Pliocene

Cubagua Formation

 Venezuela

A cusk-eel; a species of Lepophidium.

Leptolumamia[90]

Gen. et sp. nov.

Valid

Bannikov & Fraser

Lower Eocene, upper Ypresian

North-eastern Italy, Monte Bolca locality, Monte Postale site

 Italy

A new genus and species of cardinalfish (Percomorpha, Apogonidae). The type species is L. vetula.

‘Liopropoma’ sculpta[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Liopropoma

Macruronus eastmani[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene

La Meseta Formation

Antarctica (Seymour Island)
 Australia

A member of Merlucciidae, a species of Macruronus.

Malacanthus carosii[91]

Sp. nov

Valid

Carnevale

Miocene (Badenian)

Leitha Limestone

 Austria

A species of Malacanthus.

Megalichthys syndentolaminaris[64]

Sp. nov

Valid

Borgen & Nakrem

Carboniferous

 United Kingdom

A member of the family Megalichthyidae.

Melamphaes acanthifer[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Melamphaes

Nebris dioneae[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian)

Urumaco Formation

 Venezuela

A member of Sciaenidae, a species of Nebris

‘Neobythites’ auribatianus[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Neobythites

‘Neobythites’ bozzolo[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Neobythites

Neobythites huddlestoni[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (late Langhian)

Brasso Formation

 Trinidad and Tobago
(Trinidad)

A cusk-eel; a species of Neobythites.

‘Neobythites’ leonardi[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Neobythites

Neoscopelus navicularis[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Neoscopelus

Nezumia teoulerensis[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Nezumia

Notoberyx[61]

Gen. et sp. et comb. nov

Valid

Schwarzhans et al.

Early Eocene

La Meseta Formation

Antarctica (Seymour Island)
 Denmark

A slimehead. The type species is Notoberyx cionei; genus also includes Notoberyx madseni (Schwarzhans, 2007).

Occitanichthys[92]

Gen. et sp. nov

Valid

López-Arbarello & Wencker

Late Jurassic (Tithonian)

 France

A member of Semionotiformes belonging to the family Callipurbeckiidae. The type species is Occitanichthys canjuersensis.

Ogilbichthys dariensis[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (early Tortonian)

Tuira Formarion

 Panama

A viviparous brotula; a species of Ogilbichthys.

Oligoremora[93]

Gen. et sp. nov

Valid

Micklich et al.

Oligocene

 Germany

A remora. The type species is Oligoremora rhenana.

Oncorhynchus belli[94]

Sp. nov

Valid

Stearley & Smith

Miocene

Truckee Formation

 United States
( Nevada)

A species of Oncorhynchus.

Oncorhynchus rastellus[94]

Sp. nov

Valid

Stearley & Smith

Miocene

Chalk Hills Formation

 United States
( Idaho)

A species of Oncorhynchus.

Ophidion bowdenensis[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Late Pliocene

Bowden Formation

 Jamaica

A cusk-eel; a species of Ophidion.

Ophidion pauxillicauda[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Middle Pleistocene

Swan Cay Formation

 Panama

A cusk-eel; a species of Ophidion.

Ophidion sporoformis[71]

Sp. nov

Valid

Schwarzhans & Aguilera

Miocene (Tortonian)

Angostura Formation

 Ecuador

A cusk-eel; a species of Ophidion.

Ophioscion amphiamericanus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Tortonian and Messinian)

Angostura Formation
Cercado Formation

 Dominican Republic
 Ecuador

A member of Sciaenidae, a species of Ophioscion.

Ophioscion inflaticauda[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Tortonian)

Manzanilla Formation

 Trinidad and Tobago
(Trinidad)

A member of Sciaenidae, a species of Ophioscion.

Ophioscion transitivus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Burdigalian to Langhian)

Baitoa Formation
Brasso Formation

 Dominican Republic
 Trinidad and Tobago
(Trinidad)

A member of Sciaenidae, a species of Ophioscion.

‘Ophichthus’ mignolo[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Ophichthus

Orthogonikleithrus francogalliensis[95]

Sp. nov

Valid

Konwert

Late Jurassic

 France

A teleost belonging to the family Orthogonikleithridae, a species of Orthogonikleithrus.

Palaeomacrosemius[96]

Sp. nov

Valid

Ebert, Lane & Kölbl-Ebert

Late Jurassic (late Kimmeridgian to early Tithonian)

 France
 Germany

A member of Macrosemiidae. Genus includes new species P. thiollieri.

Palimphemus seymourensis[61]

Sp. nov

Valid

Schwarzhans et al.

Eocene (late Ypresian

La Meseta Formation

Antarctica (Seymour Island)

A member of Gadidae, a species of Palimphemus.

Pavarottia maiseyi[97]

Sp. nov.

Valid

Bannikov

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of Percoidei of uncertain phylogenetic placement.

Peltopleurus nitidus[98]

Sp. nov

Valid

Xu & Ma

Middle Triassic (Anisian)

 China

Pentanogmius fritschi[99]

Sp. nov

Valid

Shimada

Late Cretaceous (late Cenomanian–early Turonian)

Britton Formation

 United States
( Texas)

A member of Tselfatiiformes, a species of Pentanogmius.

‘Perca’ lactarioides[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Perca

‘Perca’ meiformis[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Perca

Phractocephalus ivy[100]

Sp. nov

Valid

Azpelicueta & Cione

Miocene (Tortonian)

Ituzaingó Formation

 Argentina

A member of the family Pimelodidae. Originally described as a species of Phractocephalus; Bogan & Agnolín (2019) transferred this species to the genus Steindachneridion.[101]

Plagioscion ultimus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Pliocene

Cubagua Formation
Springvale Formation

 Trinidad and Tobago
(Trinidad)
 Venezuela

A member of Sciaenidae, a species of Plagioscion

Pleuropholis cisnerosorum[102]

Sp. nov

Valid

Alvarado-Ortega & Brito

Late Jurassic

Sabinal Formation

 Mexico

Polycirrhus jaramilloi[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Burdigalian to Langhian)

Cantaure Formation
Castilletes Formation
Jimol Formation

 Colombia
 Venezuela

A member of Sciaenidae, a species of Polycirrhus

Polycirrhus mustus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Burdigalian to Langhian)

Cantaure Formation
Castilletes Formation
Jimol Formation

 Colombia
 Venezuela

A member of Sciaenidae, a species of Polycirrhus

Protonebris[59]

Gen. et sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Aquitanian to early Burdigalian)

Castillo Formation

 Venezuela

A member of Sciaenidae. The type species is Protonebris sanchezi

Ptychoceratodus cuyanus[103]

Sp. nov

Valid

Agnolín et al.

Late Triassic (Carnian)

Potrerillos Formation

 Argentina

A lungfish.

Rhinconichthys purgatorensis[104]

Sp. nov

Valid

Schumacher et al.

Late Cretaceous (middle Turonian)

Carlile Shale

 United States
( Colorado)

A member of Pachycormiformes, a species of Rhinconichthys.

Rhinconichthys uyenoi[104]

Sp. nov

Valid

Schumacher et al.

Late Cretaceous (Cenomanian)

Mikasa Formation

 Japan

A member of Pachycormiformes, a species of Rhinconichthys.

Saurichthys dianneae[105]

Sp. nov

Valid

Maxwell et al.

Middle Triassic (Anisian)

Vossenveld Formation

 Netherlands

Saurida trompensis[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Saurida

‘Scopelarchus’ gnocco[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Scopelarchus

Silurus spinosus[106]

Sp. nov

Valid

Kovalchuk & Ferraris

Late Miocene (late Sarmatian)

 Ukraine

A catfish; a species of Silurus.

Stellifer abbreviatus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Pliocene

Río Banano Formation

 Costa Rica

A member of Sciaenidae, a species of Stellifer.

Stellifer acerbus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Tortonian)

Manzanilla Formation

 Trinidad and Tobago
(Trinidad)

A member of Sciaenidae, a species of Stellifer.

Stellifer bicornutus[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Pliocene

Río Banano Formation

 Costa Rica

A member of Sciaenidae, a species of Stellifer.

Stellifer depressifrons[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Burdigalian to Langhian)

Cantaure Formation

 Venezuela

A member of Sciaenidae, a species of Stellifer.

Stellifer onzole[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Messinian)

Onzole Formation

 Ecuador

A member of Sciaenidae, a species of Stellifer.

Ticinolepis[107]

Gen. et 2 sp. nov

Valid

López-Arbarello et al.

Middle Triassic (Ladinian)

Besano Formation
Meride Limestone

  Switzerland

A member of Holostei of uncertain phylogenetic placement. The type species is T. longaeva; genus also includes T. crassidens.

‘Trachyrincus’ iocosus[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

Possibly a species of Trachyrincus

Turboscinetes[108]

Gen. et comb. nov

Valid

Ebert

Late Jurassic

 France
 Germany

A member of the family Pycnodontidae. The type species is "Pycnodus" egertoni Thiollière (1852).

Umbrina abbreviata[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Burdigalian to early Langhian)

Castilletes Formation

 Colombia

A member of Sciaenidae, a species of Umbrina

Umbrina bananensis[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Pliocene

Cayo Agua Formation
Río Banano Formation

 Costa Rica
 Panama

A member of Sciaenidae, a species of Umbrina

Umbrina laxa[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Burdigalian to early Langhian)

Castilletes Formation

 Colombia

A member of Sciaenidae, a species of Umbrina

Umbrina opima[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Miocene and early Pliocene

Chucunaque Formation
Cubagua Formation
Gatún Formation
Tuira Formation

 Panama
 Venezuela

A member of Sciaenidae, a species of Umbrina

Umbrina sublima[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Miocene and early Pliocene

Cayo Agua Formation
Cubagua Formation

 Panama
 Venezuela

A member of Sciaenidae, a species of Umbrina

Umbrina surda[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Late Miocene (Tortonian)

Angostura Formation

 Ecuador

A member of Sciaenidae, a species of Umbrina

Unicachichthys[109]

Gen. et sp. nov

Valid

Díaz-Cruz, Alvarado-Ortega & Carbot-Chanona

Late Cretaceous (Cenomanian)

 Mexico

A member of Aulopiformes belonging to the family Enchodontidae. The type species is Unicachichthys multidentata.

Venusichthys[110]

Gen. et sp. nov

Valid

Xu & Zhao

Middle Triassic (Anisian)

Guanling Formation

 China

A stem-neopterygian. The type species is Venusichthys comptus.

Whiteia oishii[111]

Sp. nov

Valid

Yabumoto & Brito

Triassic

 Indonesia

A coelacanth.

Wilsonichthys[112]

Gen. et sp. nov

Valid

Murray et al.

Late Cretaceous (Maastrichtian

Scollard Formation

 Canada
( Alberta)

A member of Osteoglossomorpha. The type species is Wilsonichthys aridinsulensis.

Xenotolithus semiostialis[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (late Burdigalian to early Langhian)

Brasso Formation

 Trinidad and Tobago
(Trinidad)

A member of Sciaenidae belonging to the subfamily Stelliferinae, a species of Xenotolithus.

Xenotolithus semiostialis[59]

Sp. nov

Valid

Aguilera, Schwarzhans & Béarez

Miocene (Messinian) and early Pliocene

Gros Morne Formation
Manzanilla Formation

 Trinidad and Tobago
(Trinidad)

A member of Sciaenidae belonging to the subfamily Stelliferinae, a species of Xenotolithus.

Zenion mattaccino[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Zenion

Zenion vetustus[68]

Sp. nov

Valid[69]

Lin, Nolf & Girone in Lin et al.

Eocene (Lutetian)

Aquitaine Basin

 France

A species of Zenion

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