Charmouth Mudstone Formation

Charmouth Mudstone Formation
Charmouth Mudstone Formation
Stratigraphic range: Lower Sinemurian– Lower Pliensbachian Semicostatum-lowermost Margaritatus
	Cliffs of the Charmouth Mudstone Formation at Charmouth, Dorset
Type	Formation
Unit of	Lias Group
Sub-units	Shales With Beef Member, Black Ven Marl Member, Belemnite Marl Member, Green Ammonite Member
Underlies	Dyrham Formation, Marlstone Rock Formation, unconformity with Gault (Charmouth area)
Overlies	Blue Lias Formation, Scunthorpe Mudstone
Area	Wessex Basin, Worcester Basin, East Midland Shelf
Thickness	~335 m
Lithology
Primary	Shale, mudstone
Other	Limestone, sandstone
Location
Region	England
Country	United Kingdom
Type section
Named for	Charmouth
Location	Cliff and foreshore exposures between Seven Rock Point and Golden Cap, Dorset

The Charmouth Mudstone Formation is a geological formation in England, dating to the Early Jurassic (Sinemurian–Pliensbachian).[1] It forms part of the lower Lias Group. It is most prominently exposed at its type locality in cliff section between Lyme Regis and Charmouth (alongside the underlying Blue Lias) but onshore it extends northwards to Market Weighton, Yorkshire, and in the subsurface of the East Midlands Shelf and Wessex Basin. The formation is notable for its fossils, including those of ammonites and marine reptiles and rare dinosaur remains. The formation played a prominent role in the history of early paleontology, with its Lyme Regis-Charmouth exposure being frequented by fossil collectors including Mary Anning.

Stratigraphy

Shales With Beef Member

The Shales With Beef Member is around 28–30 metres thick in the Lyme Regis-Charmouth region and predominantly consists of thinly bedded medium to dark grey mudstone, blocky calcareous pale-weathering mudstone and brown-grey organic-rich mudstones with frequent bedding parallel veins of fibrous calcite ("beef"), that are usually less than 10 centimetres thick. Several beds of nodular and tabular limestone are also present. It is the lowest unit of the formation and directly overlies the Blue Lias Formation, with the boundary being marked by a prominent bioturbated horizon. Notable persistent marker beds within the member include the laminated calcareous siltstone "Fish Bed", "Table Ledge", which consists of lens beds of limestones with mud content with nests of rhynchonellid brachiopods, the Devonshire Head and the Spittles limestones and the Birchi Nodules (which are septarian concretions) The upper boundary with the Black Ven Marl Member is marked by the prominent laterally persistent limestone Birchi Tabular Bed.[2]

Black Ven Marl Member

The Black Ven Marl Member is around 43 metres thick consists of thinly bedded dark mudstones,[3] with several laterally persistent cementstone horizons, notable horizons include the Lower and Upper Cement beds and the Stellare nodules.

Belemnite Marl Member

The Belemnite Marl Member is around 20 to 27 metres thick, and consists of interbedded pale and dark grey calcareous mudstone, with numerous belemnites, hence the name. The top of the member is marked by the Belemnite Stone Bed[4]

Green Ammonite Member

The Green Ammonite Member is up to 31 metres thick predominantly consists of medium grey mudstones, with 3 limestone horizons, Lower Limestone; Red Band, and Upper Limestone, it is conformably overlain by the Dyrham Formation in some areas,[5] but in the Charmouth area there is an erosive unconformable boundary with the much younger Early Cretaceous (Albian) aged Gault clays.

Paleobiota

Vertebrate fauna of the Charmouth Mudstone Formation

Ammonites

Ammonites of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Apoderoceras	A. cf. dunrobinense, A. subtriangulare
Arnioceras	A. semicostatum		Semicostatum Zone
Asteroceras	A. obtusum		Obtusum zone
Caenisites
Cymbites
Echioceras	E. raricostatum		Raricostatum Zone
Eoderoceras	E. bispinigerum
Gemmellaroceras	G. cf. peregrinum		Belemnite Marls
Oxynoticeras	O. williamsi
Phricodoceras	P. taylori, P. lamellosum		Belemnite Marls
Promicroceras	P. planicosta		Obtusum Zone
Radstockiceras	R. complicatum
Tragophylloceras	T. ibex, T. loscombi		Green Ammonite Member
Uptonia	U. bronni
Xipheroceras

Sauropterygia

Sauropterygia of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Archaeonectrus[6]	A. rostratus	Charmouth	Arnioceras semicostatum-Echioceras raricostatum ammonoid zone	BMNH 38525 (holotype skeleton)	A marine sauropterygian, member of the family Rhomaleosauridae
Attenborosaurus[7]	A. conybeari	Black Ven Water, Charmouth Northwest corner of Black Ven Water	Asteroceras obtusum, Arnioceras semicostatum-Echioceras raricostatum ammonoid zones	Holotype skeleton (now destroyed) BMNH 40140, partial skeleton and skull	A marine sauropterygian, early member of the family Pliosauridae. It was first identified as Plesiosaurus conybeari.	Cast of the holotype
?Plesiosaurus[8]	? Plesiosaurus sp.	Eastern point of Wear Cliffs, below Golden Cap	Prodactylioceras davoei ammonoid zone	BRSMG Ce17972a-o, articulated juvenile partial postcranial skeleton	A marine sauropterygian, type member of the family Plesiosauroidea inside Plesiosauria. A juvenile specimen that resembles those assigned to the genus Plesiosaurus. Its assignation is dubious.
Plesiosaurus[9]	P. dolichodeirus	Lyme Regis	Echioceras raricostatum ammonoid zone	Holotype (BMNH 22656) Numerous specimens referred	A marine sauropterygian, type member of the family Plesiosauroidea inside Plesiosauria. The typical plesiosaur, and one of the most common found on the formation.
Thaumatodracon[10]	T. wiedenrothi	Between Lyme Regis and Charmouth	Black Ven	NLMH 106.058, "a partial skeleton comprising a complete cranium, mandible, articulated cervical series, and indeterminate fragmentary remains"	A rhomaleosaurid

Ichthyosauria

Ichthyosauria of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Ichthyosauria indet.	Indeterminate	Lyme Regis, Charmouth		Indeterminate remains	Un-attributable to any genus
Ichthyosaurus	I. anningae[11]	Charmouth	Stonebarrow Marls Member	DONMG:1983.98 Subadult specimen	An ichthyosaurid ichthyosaur
Ichthyosaurus	Ichthyosaurus sp.	West of Westhay Water, Stonebarrow	Stonebarrow Marls Member	NHMUK R15907, partial skeleton[12]	Considered to belong to I. communis by Bennett et al. 2012, considered indeterminate within Ichthyosaurus by Lomax and Massare 2015
Leptonectes	L. moorei[13]	Seatown	Belemnite Marls Member	BMNH R14370 partial anterior skeleton with most complete skull	A leptonectid ichthyosaur
Leptonectes	L. solei[14]	Seatown	Acanthopleuroceras valdani Subzone, Tragophylloceras ibex Zone, lower Pliensbachian	NMW 91.296.2.2 "a large, isolated and incomplete forefin"	A leptonectid ichthyosaur
Temnodontosaurus	T. platyodon[15]					Temnodontosaurus platyodon.

Thalattosuchia

Thalattosuchia of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Turnersuchus[16]	T. hingleyae		Belemnite Marl Member	LRM 2021/45, a partial skeleton including cranial material, cervical, dorsal, and caudal vertebrae, ribs, right pectoral girdle, partial limb bones, and an osteoderm	An early diverging thalattosuchian.

Pterosauria

Pterosauria of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Dimorphodon[17][18][19]	D. macronyx	Charmouth Lyme Regis	Black Ven Marl Member	Tail Fragments Tibia Femur Humerus Phalanax Metacarps Semi-Complete Skeletons	A basal pterosaur and the type member of the family Dimorphodontidae.
Dimorphodontidae[19][20]	Indeterminate	Charmouth, Black Ven Lyme Regis	Black Ven Marl Member	OUM J.53070 NHMUK PV R1595, wing phalanges	A basal dimorphodontid pterosaur. Known as "Charmouth dimorphodontid" it shares resemblance with the genus Dimorphodon.
Dimorphodontidae[21]	Indeterminate	Near Seatown	Lower Pliensbachian member	Associated remains of the rostrum and mandibles	A basal dimorphodontid pterosaur. The relatively deep rostrum and extreme size dimorphism in the dentition show clear similarities to Dimorphodon, also from the Lias of Dorset. The distinctive dentition, in which the first four pairs of rostral teeth and two pairs of mandibular teeth are relatively large and fang-like, while the remaining teeth are remarkably small and short.[21]

Dinosauria

Dinosauria of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Scelidosaurus[22][23][24]	S. harrisonii	Charmouth, Black Ven Black Ven landslip	Black Ven Marl Member	NHMUK OR42068. Six osteoderms NHMUK OR42070. Phalanx. NHMUK OR42072. Large bone fragment – indet. NHMUK OR42074. Ulna (paired with NHMOR41323). NHMUK OR43062. Shaft of a large humerus, crushed proximal femur, fibula, fragmentary radius?, other large bone fragments NHMUK OR46011. Probable rib fragment – indet. BRSMG Ce12785. Partial skull and some disarticulated postcranial elements BRSMG Ce12787. Partly eroded femur and vertebra BRSMG Ce12788. cervical osteoderms. BRSMG Ce12789. Vertebra and bone fragments BRSMG Cf2781. nine middle-distal caudal vertebrae and some haemal arches with some organic (kerogenized) remains of the skin preserved. BRSMG LEGL 0004. Articulated skeleton BRSMG LEGL 0005. Articulated partial skeleton DORCM G.7542. Osteoderms DORCM G.10817. Osteoderms CAMSM X39256. Cranial: disarticulated skull and jaw elements	A basal thyreophoran, and the type member of the family Scelidosauridae. One of the best known armored dinosaurs of the lower Jurassic, also the most complete found to date, with specimens preserving even the skin and dermal armour.[25]
	Scelidosaurus sp. A[26]	Belemnite Marls at Seatown	Belemnite Marl Member, Uptonia jamesoni Zone	DORCM G.7842. Three dorsal vertebrae in articulation
	Cf. Scelidosaurus sp.[27][28][29]	Foot of Black Ven, Charmouth	Asteroceras obtusum ammonoid zone, Black Ven Marl Member	NHMUK R6704, juvenile specimen NHMUK R12019. A posterior fragment of the occiput and an articulated series of six cervical vertebrae NHMUK R10103. Indeterminate bone fragments NHMUK OR28333. Two scapulae NHMUK OR32396. Radius NHMUK OR39516. Osteoderms. NHMUK OR39517. Caudal or?sacral vertebra, small distal caudal vertebra, small distal caudal NHMUK OR39518. Haemal arch (chevron). NHMUK OR39519. Two phalanges. NHMUK OR39520. Ungual phalanx of ?pes. NHMUK OR39521. Several bone fragments. NHMUK OR40503. Tibia (proximal end). NHMUK OR41322. Femur NHMUK OR41323. Radius NHMUK OR41324. Scapular blade NHMUK OR41325. Humerus NHMUK OR41326. Fibula NHMUK OR41327. Two caudal vertebrae NHMUK OR41328. Three metatarsals NHMUK OR41329. Osteoderm NHMUK OR41330. Ischium
"Merosaurus"[22][30][31]	"M." newmani	lower cliff face, Charmouth	Black Ven Marl Member	GSM 109560, Left femur NHMUK OR 39496, Partial hind limb	A theropod, possible basal member of Tetanurae. Initially considered part of Scelidosaurus holotype, considered Tetanurae indet by Carrano 2012. It was named "Merosaurus" by Welles, Powell and Pickering in 1995.[32] But due to lack description was considered invalid.[33]
Coelophysidae[34][35]	"Megalosaurus" lydekkeri	Lyme Regis	Black Ven Marl Member	NHMUK OR 41352, isolated tooth	A theropod, possibly a member of Coelophysidae inside Neotheropoda. Invalid and non diagnostic. It was identified as Zanclodon(?) sp. b., Megalosaurus lydekkeri and reassigned as Magnosaurus(?) lydekkeri.[36] Mortimer found that was changed to "Megalosaurus" woodwardi later.[36] It was found on mostly of recent works to be non diagnostic, probably a coelophysoid.[35]
Neotheropoda[37]	Indeterminate	Near Lyme Regis	obtusum Zone, obtusum subzone	NHMUK PV R36855, Left fibula, ~ 17 cm long	A theropod, possibly a basal member of Neotheropoda. Previously misidentified as a referred specimen of the pterosaur Dimorphodon macronyx.[18] NHMUK PV R36855 differs from those of all other Early Jurassic-Late Triassic, theropods in its combination of features, but bears no discernible autapomorphies.[37] It is similar to Tawa, Dracoraptor and Liliensternus, being considered an early branching neotheropod, with a size range similar to Coelophysis bauri, representing an individual of approximately 10 kg, approximately the same size as Coelophysis bauri.[37]
Massopoda[38]	Indeterminate	lower cliff face, Charmouth	obtusum Zone, obtusum subzone	GSM 109561 pedal ungual	A sauropodomorph, possibly a member of Massopoda. While it was not associated with BMNH 39496 or GSM 109560 and was not assigned to "Merosaurus" it was classified as coming from an indeterminate theropod. Mortimer found closer resemblance with pedal unguals of basal massopods like Blikanasaurus and Jingshanosaurus.[33]

Fish

Numerous fish species are known from the Charmouth Mudstone and underlying Blue Lias, from such horizons as the "Fish Bed" of the Shales With Beef Member.

Fish of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Acrodus					A hybodont shark
Hybodus					A hybodont shark
Palidiplospinax					A synechodontiform shark
Squaloraja	S. tenuispina, S. polyspondyla				Closely related to modern chimaeras
Myriacanthus	M. paradoxus, M. granulatus		Black Ven Marl Member		A myriacanthid closely related to modern chimaeras
Dorsetichthys	D. bechei				A stem-group teleost
"Coccolepis"	"C." liassicus				A coccolepidid fish, probably does not belong to the genus
Holophagus	H. gulo				A coelacanth
Chondrosteus	C. acipenseroides				A chondrosteid acipenseriform fish, related to sturgeon and paddlefish
Oxygnathus	O. ornatus				A palaeonisciform fish
Saurorhynchus	S. brevirostris, S. anningae				A member of Saurichthyiformes
Ptycholepis	P. gracilis, P. curtus				A palaeonisciform fish
Dapedium	Spp.				A dapediiform fish
Caturus	Spp.				An amiiform fish related to bowfins
Platysiagum	P. sclerocephalum				A platysiagid fish
Furo	F. orthostomus				A member of Ionoscopiformes within Halecomorphi

Insects

Numerous species of insect are known from concretions, predominately in the Black Ven Marl Member.[39][40][41][42]

Insects of the Charmouth Mudstone Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Anglophlebia	A. gigantea			NHMUK In.51030, part and counterpart of partial forewing	Damsel-dragonfly
Archaeolepis	A. mane	Birchi Nodules, Black Ven	Shales with Beef	Isolated wing	Among the oldest known lepidopterans
Archelcana	A. durnovaria	Woodstones	Black Ven Marl Member		Elcanid orthopteran
Austaulius	A. haustrum	Monmouth Beach, Lyme Regis		NHMUK II 3103	Necrotauliid caddisfly
Brevicula	B. gradus, B. maculata	Flatstones, Woodstones		B. gradus: Holotype: NHMUK In.53993, female with ovipositor, B. maculata: Holotype: NHMUK II.3086, isolated Tegmina	Dermapterid earwig
Brochocoleus	B. maculatus				Ommatid beetle
Chrismooreia	C. michaelbehei	Charmouth	Obtusum Zone	Part and counterpart of a mostly complete specimen	Asiopterid damsel-dragonfly
Dacryoderma	B. teres	Charmouth-by-pass construction site	Obtusum Zone	Isolated tegmen	Dermapterid earwig
Dorsettia	D. laeta	Flatstones		BMNH In 59375, a male hindwing fragment	Campterophlebiid damsel-dragonfly, genus also known from China
Durnovaria	D. parallela	Flatstones			Aerophasmatid, Stem group of Phasmatodea
Elaterina	E. liassica	Lyme Regis			Click beetle
Elaterophanes	E. regius	Flatstones			Click beetle
Eoptychoptera	E. spectra	Flatstones			Ptychopterid fly, formerly referred to the genus Prodocidia
Holcoptera	H. giebeli, H. alisonae	Flatstones	Bed 83 of the Obtusum Subchronozone, Black Ven Marl Member	Complete specimen, isolated wings	Coptoclavid beetle
Hypsothemis	H. fraseri	Flatstones		BMNH In 59109, a hindwing	Campterophlebiid damsel-dragonfly, genus also known from China and Kazakhstan
Jurachorista	J. bashkuevi	Monmouth Beach, Brooki bed	Shales with Beef Member		Eomeropid scorpionfly
Lateophlebia	L. anglicanopsis	Flatstones			Campterophlebiid damsel-dragonfly
Liassocorixa	L. dorsetica	Flatstones			Corixid bug
Liassocupes	L. parvus	Flatstones			Ommatid beetle
Liassophlebia	L. pseudomagnifica	Stonebarrow		NHMUK In.64000, partial hindwing	Liassophlebiid damsel-dragonfly
Locustopsis	L. spectabilis	Flatstones			Locustopsid grasshopper
"Mesocixiella"	"M." fennahi	Flatstones			Fulgoridiid planthopper, does not belong to the genus Mesocixiella
Metaraphidia	M. confusa		Black Ven Marl Member		Snakefly
Micromacula	M. gracilis	Flatstones, Birchi Nodules			Regiatid orthopteran
Mimemala	M. giganteum	Flatstones			Schizocoleid beetle
Nannoblattina	N. petulantia	Flatstones			Mesoblattinid cockroach
Nannotanyderus	N. oliviae	Monmouth Beach, Brooki bed	Shales with Beef Member		Tanyderid fly
Neomeridium	N. trifurcum	Woodstones			Pachymeridiid Lygaeoid bug
Omma	O. liassicum	Flatstones		BMNH 59132	Ommatid beetle, genus extant
Orthophlebia	O. capillata	Flatstones		BMNH In 53924, an exoskeleton (wings & thorax)	Orthophlebiid scorpionfly
Paraprosbole	P. rotruda	Flatstones			Tettigarctid cicada
Propreocoris	P. maculatus	Woodstones			Stem-group to Ochteridae and Gelastocoridae
Protohagla	P. langi	Flatstones			Haglid cricket
Protorthophlebia	P. latipennis	Flatstones			Protorthophlebiid scorpionfly
Priscaenigma	P. obtusa	Flatstones			Snakefly
Pseudopolycentropus	P. triangularis	Woodstones	Black Ven Marl Member		Pseudopolycentropodid scorpionfly
Pterocimex	P. jacksoni	Woodstones	Black Ven Marl Member		Nepomorphan
Regiata	R. scutra	Flatstones			Regiatid orthopteran
Rossiphlebia	R. jacksoni	Flatstones		NHMUK In.53999, part and counterpart of a partial hindwing	Liassophlebiid damsel-dragonfly
Tersus	T. crowsoni	Flatstones		NHM, In. 53949	Schizophorid beetle

References

British Geological Survey. "Charmouth Mudstone Formation". BGS Lexicon of Named Rock Units. Retrieved 2018-03-08.
Gallois, R.W. (2008). The lithostratigraphy of the Shales-with-Beef Member of the Charmouth Mudstone Formation, Lower Jurassic (PDF). Ussher Society. OCLC 703253954.
"Black Ven Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
"Belemnite Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
"Green Ammonite Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
R. Owen. 1865. Part I. Sauropterygia. Monographs on the fossil Reptilia of the Liassic formations 17(75):1-40
W. J. Sollas. 1881. On a new species of Plesiosaurus (P. Conybeari) from the Lower Lias of Charmouth; with observations on P megacephalus, Stutchbury, and P. brachycephalus, Owen. Quarterly Journal of the Geological Society of London 37:440-480
G. W. Storrs. 1995. A juvenile specimen of ? Plesiosaurus sp. from the Lias (Lower Jurassic, Pliensbachian) near Charmouth, Dorset, England. Proceedings of the Dorsal Archaeological and Natural History Society 116:71-76
W. D. Conybeare. 1824. On the discovery of an almost perfect skeleton of the Plesiosaurus. Transactions of the Geological Society of London, series 2 1:381-389
Smith, Adam S.; Araújo, Ricardo (2017-07-21). "Thaumatodracon wiedenrothi, a morphometrically and stratigraphically intermediate new rhomaleosaurid plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis". Palaeontographica Abteilung A. 308 (4–6): 89–125. doi:10.1127/pala/308/2017/89. ISSN 0375-0442.
Lomax, Dean R.; Massare, Judy A. (2015-03-04). "A new species of Ichthyosaurus from the Lower Jurassic of West Dorset, England, U.K." Journal of Vertebrate Paleontology. 35 (2): e903260. doi:10.1080/02724634.2014.903260. ISSN 0272-4634. S2CID 85745787.
Bennett, S.P.; Barrett, P.M.; Collinson, M.E.; Moore-Fay, S.; Davis, P.G.; Palmer, C.P. (January 2012). "A new specimen of Ichthyosaurus communis from Dorset, UK, and its bearing on the stratigraphical range of the species". Proceedings of the Geologists' Association. 123 (1): 146–154. doi:10.1016/j.pgeola.2011.07.001.
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Lomax, Dean R.; Massare, Judy A. (December 2018). "A forefin of Leptonectes solei from the Lower Jurassic (Pliensbachian) of Dorset, UK". Proceedings of the Geologists' Association. 129 (6): 770–773. doi:10.1016/j.pgeola.2018.07.005.
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Charig AJ, Newman BH. 1992. Scelidosaurus harrisonii Owen, 1861 (Reptilia, Ornithischia): proposed replacement of inappropriate lectotype. Bulletin of Zoological Nomenclature 49: 280–283
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[BGS_Lexicon-1] British Geological Survey. "Charmouth Mudstone Formation". BGS Lexicon of Named Rock Units. Retrieved 2018-03-08.

[2] Gallois, R.W. (2008). The lithostratigraphy of the Shales-with-Beef Member of the Charmouth Mudstone Formation, Lower Jurassic (PDF). Ussher Society. OCLC 703253954.

[BGS-3] "Black Ven Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.

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