Riojasauridae

Riojasauridae is an extinct family of sauropodomorph dinosaurs from the Late Triassic Period (late Carnian to Norian Ages). It contains the genera Riojasaurus and Eucnemesaurus. The Riojasauridae is considered a stem taxon, and is defined as "the most inclusive clade containing Riojasaurus incertus but not Plateosaurus engelhardti, Massospondylus carinatus, or Anchisaurus polyzelus".[1] Geologic formations containing riojasaurid fossils include the Lower Elliot Formation of Orange Free State, South Africa (where fossils of Eucnemesaurus have been found), and the Los Colorados Formation, in La Rioja Province, Argentina (where fossils of Riojasaurus have been recovered).[1]

Riojasaurids
Temporal range: Late Triassic,
Riojasaurus skull cast
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Massopoda
Clade: Riojasauridae
Yates, 2007
Genera

Evolutionary relationships

The Riojasauridae are considered to be sauropodomorphs, but not sauropods themselves. This means that they were generally much smaller than the Sauropods of the Jurassic and Cretaceous Periods, and members of this family may not have been obligate quadrupeds, the way more derived Sauropods were. The relationships of this family within other sauropodomorphs has been considered by studies in 2007 [1] and 2020 [2] with slightly different results.

The cladogram below shows basal sauropodomorph relationships simplified after Yates, 2007. In this analysis, Riojasauridae was recovered as the earliest-diverging lineage of the clade Massopoda, and represents a sister group to the node formed by Massospondylidae, the genus Jingshanosaurus and the Anchisauria.[1]

Plateosauria
 Plateosauridae 

Unaysaurus

Plateosaurus

Massopoda
Riojasauridae

Riojasaurus

Eucnemesaurus

Massospondylidae

Massospondylus

Coloradisaurus

Lufengosaurus

Jingshanosaurus

Anchisauria

The following cladogram is from a phylogenetic analysis by Peyre de Fabrègues et al., 2020. Here, Riojasauridae was placed in a more derived position, lying closer to Sauropoda than to Massospondylidae, and appearing as the out group to the clade formed by the genus Seitaad and the Sauropodiformes.[2] The Massospondylidae and the taxa considered to be related to the family by Yates, are shown to be represent more primitive lineages. [2]

Massopoda

Plateosauravus

Ruehleia

Anchisaurus

Sarahsaurus

Massospondylidae

Xingxiulong

Jingshanosaurus

Yunnanosaurus

Riojasauridae

Eucnemesaurus

Riojasaurus

Seitaad

Sauropodiformes

References

  1. Yates, A. M. (2007). "Solving a dinosaurian puzzle: the identity of Aliwalia rex Galton". Historical Biology. 19 (1): 93–123. doi:10.1080/08912960600866953.
  2. Peyre de Fabrègues, C.; Bi, S.; Li, H.; Li, G.; Yang, L.; Xu, X. (December 2020). "A new species of early-diverging Sauropodiformes from the Lower Jurassic Fengjiahe Formation of Yunnan Province, China". Scientific Reports. 10 (1): 10961. doi:10.1038/s41598-020-67754-4.
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