Hyaenodon

Hyaenodon ("hyena-tooth") is an extinct genus of carnivorous placental mammals from extinct tribe Hyaenodontini within extinct subfamily Hyaenodontinae (in extinct family Hyaenodontidae),[19] that lived in Eurasia and North America from the middle Eocene, throughout the Oligocene, to the early Miocene.

Hyaenodon
Temporal range: Middle Eocene to Early Miocene
Mounted H. sp. skeleton, Science Museum of Minnesota
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Hyaenodonta
Superfamily: Hyaenodontoidea
Family: Hyaenodontidae
Subfamily: Hyaenodontinae
Tribe: Hyaenodontini
Leidy, 1869[1]
Genus: Hyaenodon
Laizer & Parieu, 1838
Type species
Hyaenodon leptorhynchus
Laizer and Parieu, 1838
Species
Synonyms[2]
synonyms of genus:
  • Macropterodon (Lavrov, 1999)[3]
  • Megalopterodon (Dashzeveg, 1964)
  • Microhyaenodon (Lavrov, 1999)[3]
  • Neohyaenodon (Thorpe, 1922)
  • Protohyaenodon (Stock, 1933)
  • Pseudopterodon (Schlosser, 1887)
  • Taxotherium (Blainville, 1841)
synonyms of species:
  • H. brachyrhynchus:
    • Canis brachyrhynchus (Blainville, 1841)
    • Hyaenodon cuvieri (Pictet, 1853)[4]
    • Hyaenodon leptorhynchus (Dujardin, 1840)[5]
    • Hyaenodon parisiensis (Laurillard, 1845)[6]
    • Hyaenodon vulpinus (Gervais, 1873)
    • Nasua parisiensis (Blainville, 1841)
    • Pterodon brachyrhynchus (Pomel, 1846)[7]
    • Pterodon cuvieri (Pomel, 1846)
    • Taxotherium parisiense (Blainville, 1841)
  • H. brevirostrus:
    • Hyaenodon brevirostris (Joeckel, 1997)[8]
    • Protohyaenodon brevirostrus (Mellett, 1977)
  • H. chunkhtensis:
    • Microhyaenodon chunkhtensis (Lavrov, 1999)
  • H. crucians:
    • Hyaenodon leptocephalus (Scott, 1888)[9]
    • Hyaenodon minutus (Douglass, 1902)
    • Hyaenodon paucidens (Osborn & Wortman, 1894)[10]
    • Protohyaenodon crucians (Mellett, 1977)
    • Prtohyaenodon crucians (Lavrov, 1999)
    • Pseudopterodon minutus (Douglass, 1902)
  • H. dubius:
    • Hyaenodon aymardi (Filhol, 1882)[11]
  • H. exiguus:
    • Hyaenodon exigus (Lavrov, 1999)
    • Pterodon exiguum (Gervais, 1873)
  • H. filholi:
    • Hyaenodon compressus (Filhol, 1876)
    • Hyaenodon vulpinus (Filhol, 1876)
    • Microhyaenodon filholi (Lavrov, 1999)
    • Pseudopterodon ganodus (Schlosser, 1887)
  • H. gervaisi:
    • Hyaenodon ambiguus (Martin, 1906)
  • H. gigas:
    • Macropterodon zelenovi (Lavrov, 1999)
    • Neohyaenodon gigas (Lavrov, 1999)
  • H. heberti:
    • Hyaenodon arnaudi (Depéret, 1917)[12]
  • H. horridus:
    • Hyaenodon cruentus (Leidy, 1853)
    • Neohyaenodon horridus (Thorpe, 1922)
    • Neohyaenodon semseyi (Kretzoi, 1941)[13]
  • H. incertus:
    • Neohyaenodon incertus (Lavrov, 1999)
    • Pterodon exploratus (Dashzeveg, 1985)
  • H. leptorhynchus:
    • Canis leptorhynchus (Blainville, 1841)
    • Hyaenodon bavaricus (Dehm, 1935)[14]
    • Hyaenodon cayluxi (Filhol, 1876)
    • Hyaenodon martini (Depéret, 1917)
    • Hyaenodon milloquensis (Martin, 1906)
    • Pterodon leptorhynchus (Pomel, 1846)
  • H. macrocephalus:
    • Neohyaenodon macrocephalus (Lavrov, 1999)
  • H. megaloides:
    • Neohyaenodon megaloides (Mellett, 1977)
  • H. microdon:
    • Microhyaenodon microdon (Lavrov, 1999)
    • Protohyaenodon microdon (Mellett, 1977)
  • H. milvinus:
    • Neohyaenodon milvinus (Lavrov, 1999)
  • H. minor:
    • Hyaenodon aimi (Cooper, 1926)[15]
  • H. mongoliensis:
    • Epipterodon mongoliensis (Lavrov, 1999)
    • Megalopterodon mongoliensis (Dashzeveg, 1964)
    • Neohyaenodon mongoliensis (Morlo & Nagel, 2006)[16]
    • Pterodon mongoliensis (Van Valen, 1967)[17]
  • H. montanus:
    • Protohyaenodon montanus
    • Neohyaenodon montanus (Mellett, 1977)
  • H. mustelinus:
    • Protohyaenodon mustelinus (Scott, 1894)
  • H. pervagus:
    • Hyaenodon neimongoliensis (Huang & Zhu, 2002)[18]
  • H. pumilus:
    • Microhyaenodon pumilus (Lavrov, 1999)
  • H. raineyi:
    • Microhyaenodon raineyi (Lavrov, 1999)
    • Protohyaenodon raineyi (Gustafson, 1986)
  • H. requieni:
    • Hyaenodon heberti euzetensis (Depéret, 1917)
    • Pterodon requieni (Gervais, 1846)
  • H. venturae:
    • Hyaenodon exiguus (Stock, 1933)
    • Microhyaenodon venturae (Lavrov, 1999)
    • Protohyaenodon exiguus (Stock, 1933)
    • Protohyaenodon venturae (Mellett, 1977)
  • H. vetus:
    • Neohyaenodon vetus (Mellett, 1977)
    • Pterodon californicus (Stock, 1933)
  • H. yuanchuensis:
    • Hyaenodon yuanchüensis (Young, 1937)

Description

Skull of Hyaenodon horridus
Life reconstruction of H. horridus

Typical of early carnivorous mammals, individuals of Hyaenodon had a very massive skull, but a small brain. The skull is long with a narrow snout - much larger in relation to the length of the skull than in canine carnivores, for instance. The neck was shorter than the skull, while the body was long and robust and terminated in a long tail. Compared to the larger (but not closely related) Hyainailouros, the dentition of Hyaenodon was geared more towards shearing meat and less towards bone crushing.[20]

Some species of this genus were among the largest terrestrial carnivorous mammals of their time; others were only of the size of a marten. Remains of many species are known from North America, Europe, and Asia.[21] The average weight of adult or subadult H. horridus, the largest North American species, is estimated to about 40 kg (88 lb) and may not have exceeded 60 kg (130 lb). H. gigas, the largest Hyaenodon species, was much larger, being 378 kg (833 lb) and around 10 feet (3.0 m).[20] H. crucians from the early Oligocene of North America is estimated to only 10 to 25 kg (22 to 55 lb). H. microdon and H. mustelinus from the late Eocene of North America were even smaller and weighed probably about 5 kg (11 lb).[22]

Tooth eruption

Studies on juvenile Hyaenodon specimens show that the animal had a very unusual system of tooth replacement. Juveniles took about 3–4 years to complete the final stage of eruption, implying a long adolescent phase. In North American forms, the first upper premolar erupts before the first upper molar, while European forms show an earlier eruption of the first upper molar.[23]

Paleoecology

The various species of Hyaenodon competed with each other and with other hyaenodont genera (including Sinopa, Dissopsalis, and Hyainailurus), and played important roles as predators in ecological communities as late as the Miocene in Asia and preyed on a variety of prey species such as primitive horses like Mesohippus, Brontotheres, early camels, oreodonts and even primitive rhinos.[20] Species of Hyaenodon have been shown to have successfully preyed on other large carnivores of their time, including a nimravid ("false sabertooth cat"), according to analysis of tooth puncture marks on a fossil Dinictis skull found in North Dakota.[24][25]

In North America the last Hyaenodon, in the form of H. brevirostrus, disappeared in the late Oligocene.[26] In Europe, they had already vanished earlier in the Oligocene.[21]

Classification and phylogeny

Reconstruction of Hyaenodon by Heinrich Harder (around 1920.)
Reconstruction of H. horridus and Leptomeryx evansi by W. B. Scott (1913.)

Taxonomy

  • Tribe: †Hyaenodontini (Leidy, 1869)
    • Genus: †Hyaenodon (Laizer & Parieu, 1838)
      • Hyaenodon brachyrhynchus (Blainville, 1841)[27]
      • Hyaenodon chunkhtensis (Dashzeveg, 1985)[28]
      • Hyaenodon dubius (Filhol, 1873)[29]
      • Hyaenodon eminus (Matthew & Granger, 1925)[30]
      • Hyaenodon exiguus (Gervais, 1873)[31]
      • Hyaenodon filholi (Schlosser, 1887)[32]
      • Hyaenodon gervaisi (Martin, 1906)[33]
      • Hyaenodon heberti (Europe, 41-33.9 mya) (Filhol, 1876)[34]
      • Hyaenodon leptorhynchus (Laizer & Parieu, 1838)[35]
      • Hyaenodon minor (Lange-Badré, 1979)[36]
      • Hyaenodon pervagus (Matthew & Granger, 1924)[37]
      • Hyaenodon pumilus (Lavrov, 2019[38]
      • Hyaenodon requieni (Gervais, 1846)[39]
      • Hyaenodon rossignoli (Lange-Badré, 1979)
      • Hyaenodon weilini (China, 23-16,9 mya) (Wang, 2005)[21]
      • Hyaenodon yuanchuensis (Young, 1937)[40]
      • Subgenus: †Neohyaenodon (paraphyletic subgenus) (Thorpe, 1922)[41]
        • Hyaenodon gigas (Dashzeveg, 1985)
        • Hyaenodon horridus (Leidy, 1853)[42]
        • Hyaenodon incertus (Dashzeveg, 1985)
        • Hyaenodon macrocephalus (Lavrov, 1999)[3]
        • Hyaenodon megaloides (Mellett, 1977)[43]
        • Hyaenodon milvinus (Lavrov, 1999)[3]
        • Hyaenodon mongoliensis (Dashzeveg, 1964)[44]
        • Hyaenodon montanus (Douglass, 1902)[45]
        • Hyaenodon vetus (Stock, 1933)[46]
      • Subgenus: †Protohyaenodon (paraphyletic subgenus) (Stock, 1933)
        • Hyaenodon brevirostrus (Macdonald, 1970)[47]
        • Hyaenodon crucians (Leidy, 1853)
        • Hyaenodon microdon (Mellett, 1977)
        • Hyaenodon mustelinus (North America, 38-30 mya) (Scott, 1894)[48]
        • Hyaenodon raineyi (Gustafson, 1986)[49]
        • Hyaenodon venturae (Mellett, 1977)

Phylogeny

The phylogenetic relationships of genus Hyaenodon are shown in the following cladogram[3][50][51][52][53][54]

 Hyaenodontidae 

Paracynohyaenodon

Quercytherium

Cynohyaenodon lautricensis

Cynohyaenodon cayluxi

Cynohyaenodon ruetimeyeri

Cynohyaenodon trux

Cynohyaenodon

Cynohyaenodon smithae

Boritia

 ? 

Protoproviverra

 ? 

Praecodens

Leonhardtina

Preregidens

Cartierodon

Prodissopsalis

 ? 

Paenoxyaenoides

Eurotherium theriodis

Eurotherium mapplethorpei

Eurotherium

Eurotherium matthesi

Alienetherium

Matthodon

Oxyaenoidinae

 ? 

Neosinopa

 Hyaenodontinae 
 ? 

Consobrinus

Propterodon morrisi

Propterodon sp. (ZIN no. 34494)

Propterodon paganensis

Propterodon tongi

Propterodon witteri

Epipterodontini

 Hyaenodontini 
 Hyaenodon 

Hyaenodon chunkhtensis

Hyaenodon eminus

 Hyaenodon venturae 

 (†Protohyaenodon) 

Hyaenodon pumilus

Hyaenodon filholi

 Hyaenodon microdon 

 Hyaenodon raineyi 

 Hyaenodon mustelinus 

 Hyaenodon brevirostrus 

 Hyaenodon crucians 

Hyaenodon yuanchuensis

Hyaenodon brachyrhynchus

Hyaenodon gervaisi

Hyaenodon requieni

Hyaenodon minor

Hyaenodon dubius

Hyaenodon leptorhynchus

Hyaenodon exiguus

Hyaenodon sp. (Ergiliyn Dzo)

Hyaenodon pervagus

Hyaenodon rossignoli

Hyaenodon horridus

Hyaenodon megaloides

Hyaenodon montanus

Hyaenodon vetus

Hyaenodon macrocephalus

Hyaenodon milvinus

Hyaenodon gigas

Hyaenodon incertus

Hyaenodon mongoliensis

Hyaenodon heberti

Hyaenodon weilini

Cynohyaenodon/Quercytherium clade
Leonhardtina clade
Eurotherium clade
Matthodon clade
Oxyaenoides clade
Propterodon

See also


References

  1. Leidy J. (1869.) "On the extinct Mammalia of Dakota and Nebraska: including an account of some allied forms from other localities, together with a synopsis of the mammalian remains of North America." Journal of the Academy of Natural Sciences Philadelphia 7: 1–472.
  2. J. Alroy (2002.) "Synonymies and reidentifications of North American fossil mammals."
  3. A. V. Lavrov (1999.) "Adaptive Radiation of Hyaenodontinae (Creodonta, Hyaenodontidae) of Asia." in 6th Congress of the Theriological Society, Moscow, April 13–16, p. 138 [in Russian].
  4. F. J. Pictet (1853.) "Traité de Paléontologie." I (2e edit.):584 p. + atlas 110 pl.
  5. F. Dujardin (1840.) "Note sur une tête fossile Hyaenodon trouvée au bord du Tarn, près de Rabastens." Comptes-Rendus de l'Académie des Sciences de Paris, 10:134-135
  6. Ch. Laurillard (1845.) "Hyaenodon." in: d'Orbigny: "Dictionnaire Universel d'Histoire naturelle 1-6": 767-769, Renard, Martinet édit., Paris
  7. A. Pomel (1846.) "Note sur le Pterodon, genre voisin des Dasyures dont plusieurs espèces ont été trouvées dans les terrains tertiaires." Bulletin de la Société Géologique de France, 4(2):385-393
  8. R. M. Joeckel, H. W. Bond and G. W. Kabalka (1997.) "Internal Anatomy of the Snout and Paranasal Sinuses of Hyaenodon (Mammalia, Creodonta)" Journal of Vertebrate Paleontology, Vol. 17, No. 2, pp. 440-446
  9. W. B. Scott (1888.) "On some new and little know creodonts." Journal of the Academy of Natural Sciences of Philadelphia 9:155-185
  10. H. F. Osborn and J. L. Wortman (1894.) "Fossil mammals of the Lower Miocene White River beds. Collection of 1892." Bulletin of the American Museum of Natural History 6(7):199-228
  11. H. Filhol (1882.) "Étude des Mammifères de Ronzon (Haute-Loire)." Ann. Sci. géol., Paris, 12(3):270 p., 25 pl.
  12. Depéret C. (1917.) "Monographie de la faune de mammiféres fossiles du Ludien inférieur d'Euget-les-Bains (Gard)." Ann. Univ. Lyon (N. S.), Div. 1, 40, 1-288.
  13. Kretzoi, M. (1941.) "Ausländische Säugetierfossilien der Ungarischen Museen. (1-4)" Földtani Közlöny, vol. 71, nos. 4-6, pp. 1-6
  14. R. Dehm (1935.) "Über tertiäre Spaltenfüllungen im Fränkischen und Schwäbischen Jura [On Tertiary fissure fillings in the Franconian and Swabian Jura."] Abhandlungen der Bayerischen Akademie der Wissenschaften Mathematisch-naturwissenschaftliche Abteilung, Neue Folge 29:1-86
  15. Forster Cooper, C. (1926.) "Hyænodon aimi, sp. n., and a note on the occurrence of Anthracotherium minus from the Headon Beds at Hordle." Annals and Magazine of Natural History (9)18(106):370–373
  16. Michael Morlo, Doris Nagel (2006.) "New remains of Hyaenodontidae (Creodonta, Mammalia) from the Oligocene of Central Mongolia." Annales de Paléontologie 92(3):305-321
  17. L. Van Valen (1967.) "New Paleocene insectivores and insectivore classification." Bulletin of the American Museum of Natural History 135(5):217-284
  18. Xueshi Huang, Ba Zhu (2002.) "Creodont (Mammalia) remains from the Early Oligocene of Ulantatal, Nei Mongol." Vertebrata PalAsiatica 40(1):17-22
  19. Malcolm C. McKenna, Susan K. Bell (1997.) "Classification of Mammals: Above the Species Level", Columbia University Press, New York, 631 pages. Hyaenodon
  20. Wang, Xiaoming; Tedford, Richard H. (2008). Dogs: Their Fossil Relatives and Evolutionary History. New York: Columbia University Press. p. 1. doi:10.7312/wang13528. ISBN 978-0-231-13528-3. JSTOR 10.7312/wang13528.
  21. X. Wang, Z. Qiu and B. Wang (2005.) "Hyaenodonts and carnivorans from the early Oligocene to early Miocene of the Xianshuihe Formation, Lanzhou Basin, Gansu Province, China." Palaeontologia Electronica 8(1):1-14
  22. Egi, Naoko (2001). "Body mass estimates in extinct mammals from limb bone dimensions: the case of North American hyaenodontids". Palaeontology. 44 (3): 497–528. doi:10.1111/1475-4983.00189. S2CID 128832577.
  23. Bastl, Katharina Anna (2013). "First evidence of the tooth eruption sequence of the upper jaw in Hyaenodon (Hyaenodontidae, Mammalia) and new information on the ontogenetic development of its dentition". Paläontologische Zeitschrift. 88 (4): 481–494. doi:10.1007/s12542-013-0207-z. S2CID 85304920.
  24. John W. Hoganson and Jeff Person (2010.) "Tooth puncture marks on a skull of Dinictis (Nimravidae) from the Oligocene Brule Formation of Northe Dakota attributed to predation by Hyaenodon (Hyaenodontidae)", North Dakota Geological Survey
  25. John W. Hoganson and Jeff Person (2011.) "Tooth puncture marks on a 30 million year old Dinictis skull.", Geo News, p. 12-17
  26. Van Valkenburgh, Blaire (1994). "Extinction and replacement among predatory mammals in the North American late Eocene and Oligocene: Tracking a paleoguild over twelve million years". Historical Biology. 8 (1–4): 129–150. doi:10.1080/10292389409380474. Retrieved 12 April 2022.
  27. Henri Marie Ducrotay de Blainville (1841.) "Ostéographie ou description iconographique comparée du squelette et du système dentaire des mammifères récents et fossiles." Tome 2: Secondates et Subursus, 123 p.; Viverras, 100 p. et atlas, 117 pl. Baillėte édit. Paris.
  28. Dashzeveg D. (1985.) "Nouveau Hyaenodontinae (Creodonta, Mammalia) du Paléogène de Mongolie." Annales de Paléontologie 71:223–256
  29. Filhol, H. (1873.) "Sur les Vertébrés fossiles trouvés dans les dépôts de phosphate de chaux du Quercy." Bull. Soc. Pholomath. Paris (6) 10, 85-89.
  30. Matthew W. D. & Granger W. (1925.) "New creodonts and rodents from the Ardyn Obo Formation of Mongolia." American Museum Novitates 193:1–11.
  31. Gervais P. (1873.) "Mammifères dont les ossements accompagnent les dépôts de chaux phosphatée des départements du Tarn-et.Garonne et du Lot." Journal de Zoologie, Paris, 2:356-380
  32. M. Schlosser (1887.) "Die Affen, Lemuren, Chiropteren, Insectivoren, Marsupialier, Creodonten und Carnivoren des Europaischen Tertiars." Beitrage zur Paleontologie Osterreich-Ungarns un des Orients 6:1-224
  33. R. Martin (1906.) "Revision der obereocænen und unteroligocænen Creodonten Europas." Rev. Suisse Zool., 14, (3), pp. 405-500
  34. H. Filhol (1876.) "Recherches sur les phospohorites du Quercy. Étude des fossiles qi'on y rencontre et spécialement des Mammifères." Annales des Sciences Géologiques, Paris, 7(7):220 p., pl. 11-36; 1877:art. 1, 340 p., 28 pl.
  35. Laizer, L. D. and Parieu, D. (1838.) "Description et détermination d'une mâchoire fossile appartenant à un mammifère jusqu'à présent inconnu, Hyaenodon leptorhynchus." Comptes Rendus de l'Académie des Sciences Paris, 7:442
  36. Lange-Badré B. (1979.) "Les créodontes (Mammalia) d'Europe occidentale de l'Éocène supérieur à l'Oligocène supérieur." Mémoires du Muséum National d'Histoire Naturelle 42: 1–249
  37. W. D. Matthew and W. Granger (1924.) "New Carnivora from the Tertiary of Mongolia." American Museum Novitates 104:1-9
  38. A. V. Lavrov (2019). "New material on small hyenodons Hyaenodontinae, Creodonta) from the Paleogene of Mongolia". Paleontological Journal. 53 (4): 418–431. doi:10.1134/S0031030119040063. S2CID 201654889.
  39. Gervais P. (1846.) "Mémoire sur quelques Mammifères fossiles du Vaucluse." Comptes rendus hebdomadaires des séances de l'Académie des sciences, Paris, T. 22, pp. 845-846.
  40. C. Young (1937.) "An early Tertiary vertebrate fauna from Yuanchü." Bulletin of the geological society of China 17(3-4):413-438
  41. M. R. Thorpe (1922.) "A new genus of Oligocene Hyaenodontidae." American Journal of Science 3(16):277-287
  42. Leidy, J. (1853.) "Remarks on a collection of fossil Mammalia from Nebraska." Proceedings of the Academy of Natural Sciences, Philadelphia, 6:392-394.
  43. J. S. Mellett (1977.) "Paleobiology of North American Hyaenodon (Mammalia, Creodonta)." Contributions to Vertebrate Evolution 1:1-134
  44. D. Dashzeveg (1964.) "On two Oligocene Hyaenodontidae from Erghilyin-Dzo (Mongolian People's Republic)." Acta Palaeontologica Polonica 9(2):263-274
  45. E. Douglass (1902.) "Fossil Mammalia of the White River beds of Montana." Transactions of the American Philosophical Society 20:237-279
  46. C. Stock (1933.) "Hyaenodontidae of the Upper Eocene of California." Proceedings of the National Academy of Sciences 19(4):434-440
  47. J. R. Macdonald (1970.) "Review of the Miocene Wounded Knee faunas of southwestern South Dakota." Bulletin of the Los Angeles County Museum of Natural History, Science 8:165-82
  48. William Berryman Scott (1894.) "The osteology of Hyaenodon" Journal of Academy of Natural Sciences, Philadelphia (2), 9, 291-323
  49. E. P. Gustafson (1986.) "Carnivorous mammals of the Late Eocene and Early Oligocene of Trans-Pecos Texas." Texas Memorial Museum Bulletin 33:1-66
  50. Borths, Matthew R; Stevens, Nancy J (2017). "Deciduous dentition and dental eruption of Hyainailouroidea (Hyaenodonta, "Creodonta," Placentalia, Mammalia)". Palaeontologia Electronica. 20 (3): 55A. doi:10.26879/776.
  51. Matthew R. Borths; Nancy J. Stevens (2019). "Simbakubwa kutokaafrika, gen. et sp. nov. (Hyainailourinae, Hyaenodonta, 'Creodonta,' Mammalia), a gigantic carnivore from the earliest Miocene of Kenya". Journal of Vertebrate Paleontology. 39 (1): e1570222. Bibcode:2019JVPal..39E0222B. doi:10.1080/02724634.2019.1570222. S2CID 145972918.
  52. Floréal Solé; Bernard Marandat; Fabrice Lihoreau (2020). "The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian". Geodiversitas. 42 (13): 185–214. doi:10.5252/geodiversitas2020v42a13. S2CID 219585388.
  53. Solé, F.; Morlo, M.; Schaal, T.; Lehmann, T. (2021). "New hyaenodonts (Mammalia) from the late Ypresian locality of Prémontré (France) support a radiation of the hyaenodonts in Europe already at the end of the early Eocene". Geobios. 66–67: 119–141. Bibcode:2021Geobi..66..119S. doi:10.1016/j.geobios.2021.02.004. S2CID 234848856.
  54. Averianov, Alexander; Obraztsova, Ekaterina; Danilov, Igor; Jin, Jian-Hua (2023). "A new hypercarnivorous hyaenodont from the Eocene of South China". Frontiers in Ecology and Evolution. 11. doi:10.3389/fevo.2023.1076819. ISSN 2296-701X.
This article is issued from Wikipedia. The text is licensed under Creative Commons - Attribution - Sharealike. Additional terms may apply for the media files.